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bilobed speedwell, sawtooth speedwell, twolobed speedwell

spike speedwell, véronique en épi

Habit Annuals. Perennials.
Stems

erect, (7–)10–20(–30) cm, eglandular- and/or glandular-hairy.

ascending, usually with 4–10 nodes, 5–45(–60) cm, proximally eglandular- or glandular-hairy, sometimes glabrous, distally eglandular- and, usually, glandular-hairy.

Leaves

blade lanceolate or elliptic, (6–)12–28(–35) × 3–12(–20) mm, base cuneate, margins serrate, teeth 4–7(–12) per side, apex acute, surfaces sparsely glandular-hairy or glabrate.

blade ovate-oblong to narrowly oblong-lanceolate, (30–)40–70(–80) × 5–20 mm, base long-cuneate, margins shallowly crenate to shallowly serrate to subentire, apex obtuse to rounded, surfaces densely glandular- or eglandular-hairy, rarely glabrate.

Racemes

1–8, terminal, usually reduced and replaced by 2 pseudodichotomous racemes with additional axillary racemes, 80–150 mm, (5–)10–25(–35)-flowered, axis eglandular- and/or glandular-hairy;

bracts lanceolate, 9–12 mm, proximals with margins serrate, distals sometimes entire.

1(–7), terminal, sometimes with lateral ones, (50–)70–130(–170) mm, (50–)100–300-flowered, axis eglandular- and glandular-hairy;

bracts oblong to lanceolate, 3–5 mm.

Pedicels

erect-patent to spreading proximally, deflexed distally, (3–)6–9(–12) mm, shorter than subtending bract, densely glandular- and eglandular-hairy.

suberect to patent, 0.5–1(–2) mm, shorter than subtending bract, eglandular- and glandular-hairy.

Flowers

calyx lobes (5–)6–8.5(–10) mm, apex acuminate-acute, pairs connate 0.8–1.1 mm, glandular- and eglandular-hairy, rarely sparsely hairy;

corolla mostly intense blue with white center, 4–6 mm diam.;

stamens 2.5 mm;

style (0.7–)1–1.3(–1.7) mm.

calyx lobes 2–3 mm, ciliate, apex obtuse, glandular-hairy, rarely eglandular-hairy;

corolla blue, campanulate, longer than wide, 5–6 mm diam., lobes 3–4.5 × 1.5–2 mm;

stamens 5 mm;

style 8 mm.

Capsules

slightly compressed in cross section, 3.5–5 × (4–)5–8 mm, apex emarginate, sinus angle 30–45(–60)°, densely eglandular- and glandular-hairy, rarely eglandular-hairy.

± compressed in cross section, broadly ovoid, 2–3 × 2–3 mm, apex obtuse, densely glandular-puberulent.

Seeds

4–10, yellowish brown, oblong to ovoid, cymbiform, 1.5–2.5 × 0.8–1.5 mm, 0.8 mm thick, cristate, dorsally reticulate-verrucate.

3–40, light brown, ellipsoid, flat, 0.6–1.2 × 0.5–0.8 mm, 0.2–0.4 mm thick, smooth.

2n

= 18, 42 (Asia).

= 34, 68 (Europe).

Veronica argute-serrata

Veronica spicata

Phenology Flowering Apr–Jun. Flowering Jun–Sep.
Habitat Bare soils, oak and juniper forests, steppes, alpine meadows, fields, stony slopes, streams, gardens, ruins, ruderal places, calcareous and siliceous soils. Dry grasslands.
Elevation (600–)900–3000 m. ((2000–)3000–9800 ft.) 0–700 m. (0–2300 ft.)
Distribution
from FNA
CO; ID; IL; KS; MT; NE; NM; NV; NY; OR; UT; WA; WY; BC; Eurasia [Introduced in North America]
[WildflowerSearch map]
from FNA
CT; NH; NY; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

Veronica argute-serrata has been recorded mostly under the name V. biloba and occasionally under V. campylopoda Boissier. Most of the records for V. biloba likely belong under this name. Veronica biloba is also found in the flora area, sometimes mixed with V. argute-serrata; V. campylopoda has not been recorded from the flora area. A chromosome count of 2n = 18 (C. R. Bell 1965) for American material is suspicious because 2n = 42 has been reported multiple times in Turkish plants, and a base chromosome number of × = 9 is not known in related species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica spicata is widely distributed in horticulture and a multitude of cultivars is available. Some specimens may also be derived from a related species, V. barrelieri Schott ex Roemer & Schultes, differentiated from V. spicata by glabrous calyx lobes, and in its typical variety, by eglandular pubescence. Another closely related species, V. incana Linnaeus, has a dense white woolly indumentum and is native to northern Asia and eastern Europe; it has not escaped from cultivation in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 308.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. bornmuelleri
Name authority Regel & Schmalhausen: Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 626. (1878) — (as argute serrata) Linnaeus: Sp. Pl. 1: 10. (1753)
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