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bilobed speedwell, sawtooth speedwell, twolobed speedwell

gray field speedwell, gray speedwell

Habit Annuals. Annuals.
Stems

erect, (7–)10–20(–30) cm, eglandular- and/or glandular-hairy.

creeping to decumbent, 5–20(–40) cm, eglandular-hairy.

Leaves

blade lanceolate or elliptic, (6–)12–28(–35) × 3–12(–20) mm, base cuneate, margins serrate, teeth 4–7(–12) per side, apex acute, surfaces sparsely glandular-hairy or glabrate.

blade suborbiculate to ovate, rarely oblong-ovate, (3–)6–11(–14) × (3–)4–9(–10) mm, base truncate, margins serrate to dentate, teeth 2–4 per side, apices usually rounded, apex acute to obtuse, abaxial surface densely hairy, adaxial sparsely hairy to glabrate.

Racemes

1–8, terminal, usually reduced and replaced by 2 pseudodichotomous racemes with additional axillary racemes, 80–150 mm, (5–)10–25(–35)-flowered, axis eglandular- and/or glandular-hairy;

bracts lanceolate, 9–12 mm, proximals with margins serrate, distals sometimes entire.

1–4, terminal, 50–200(–400) mm, 5–20-flowered, axis sparsely to densely hairy or glabrate;

bracts suborbiculate to ovate, (3–)6–11(–14) mm.

Pedicels

erect-patent to spreading proximally, deflexed distally, (3–)6–9(–12) mm, shorter than subtending bract, densely glandular- and eglandular-hairy.

usually semicircularly recurved, (3–)6–13(–15) mm, 0.5–1.5 times subtending bract, densely eglandular-hairy.

Flowers

calyx lobes (5–)6–8.5(–10) mm, apex acuminate-acute, pairs connate 0.8–1.1 mm, glandular- and eglandular-hairy, rarely sparsely hairy;

corolla mostly intense blue with white center, 4–6 mm diam.;

stamens 2.5 mm;

style (0.7–)1–1.3(–1.7) mm.

calyx lobes broadly ovate, (3–)4–6(–7) mm, 2.6–3.8 mm wide, apex acute, eglandular-hairy, rarely glandular-hairy;

corolla intense dark to bright blue, rarely pale lilac or white, with darker nerves, abaxial 1/2 often brighter to whitish, (3–)4–7(–8) mm diam.;

stamens 0.5 mm;

style (0.5–)1–1.6(–1.8) mm.

Capsules

slightly compressed in cross section, 3.5–5 × (4–)5–8 mm, apex emarginate, sinus angle 30–45(–60)°, densely eglandular- and glandular-hairy, rarely eglandular-hairy.

± round in cross section, obcordiform, (2.5–)3–4(–4.5) × (3.5–)4–6(–6.3) mm, apex emarginate, sinus angle 20–60(–80)°, densely eglandular-hairy, sometimes also glandular-puberulent, rarely glabrous.

Seeds

4–10, yellowish brown, oblong to ovoid, cymbiform, 1.5–2.5 × 0.8–1.5 mm, 0.8 mm thick, cristate, dorsally reticulate-verrucate.

(7–)16–24(–30), pale yellow, ellipsoid to ovoid, deeply cymbiform, 0.9–1.6 × (0.5–)0.8–1.3 mm, 0.4–0.8 mm thick, rugose-cristate.

2n

= 18, 42 (Asia).

= 14 (Eurasia).

Veronica argute-serrata

Veronica polita

Phenology Flowering Apr–Jun. Flowering (Feb–)Mar–Jun(–Jul).
Habitat Bare soils, oak and juniper forests, steppes, alpine meadows, fields, stony slopes, streams, gardens, ruins, ruderal places, calcareous and siliceous soils. Fields, ruderal places, calcareous soils, lawns.
Elevation (600–)900–3000 m. ((2000–)3000–9800 ft.) 0–600 m. (0–2000 ft.)
Distribution
from FNA
CO; ID; IL; KS; MT; NE; NM; NV; NY; OR; UT; WA; WY; BC; Eurasia [Introduced in North America]
[WildflowerSearch map]
from FNA
AL; AR; AZ; CO; CT; DC; FL; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; NC; NE; NY; OH; OK; PA; TN; TX; VA; WI; WV; MB; NB; ON; Eurasia; nw Africa (Algeria, Morocco) [Introduced in North America; introduced also in Mexico, s South America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica argute-serrata has been recorded mostly under the name V. biloba and occasionally under V. campylopoda Boissier. Most of the records for V. biloba likely belong under this name. Veronica biloba is also found in the flora area, sometimes mixed with V. argute-serrata; V. campylopoda has not been recorded from the flora area. A chromosome count of 2n = 18 (C. R. Bell 1965) for American material is suspicious because 2n = 42 has been reported multiple times in Turkish plants, and a base chromosome number of × = 9 is not known in related species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Veronica polita are similar to those of the more frequent V. persica and are probably frequently overlooked.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 319.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. bornmuelleri
Name authority Regel & Schmalhausen: Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 626. (1878) — (as argute serrata) Fries: Novit. Fl. Svec., 63. (1819)
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