Veronica argute-serrata |
Veronica officinalis |
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bilobed speedwell, sawtooth speedwell, twolobed speedwell |
common gypsyweed, common speedwell, heath speedwell, herbal speedwell, Paul's betony, véronique officinale |
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Habit | Annuals. | Perennials. |
Stems | erect, (7–)10–20(–30) cm, eglandular- and/or glandular-hairy. |
creeping, decumbent to ascending distally, 10–40(–50) cm, densely eglandular-hairy, rarely glabrate or glabrous. |
Leaves | blade lanceolate or elliptic, (6–)12–28(–35) × 3–12(–20) mm, base cuneate, margins serrate, teeth 4–7(–12) per side, apex acute, surfaces sparsely glandular-hairy or glabrate. |
blade ovate, ovate-oblong, or obovate, (8–)20–35(–50) × (3–)10–20(–30) mm, 1.5–2 times as long as wide, base attenuate to cuneate or obtuse, margins dentate, denticulate, or serrulate, apex obtuse to rounded, surfaces sparsely to densely eglandular-hairy, rarely glabrate. |
Racemes | 1–8, terminal, usually reduced and replaced by 2 pseudodichotomous racemes with additional axillary racemes, 80–150 mm, (5–)10–25(–35)-flowered, axis eglandular- and/or glandular-hairy; bracts lanceolate, 9–12 mm, proximals with margins serrate, distals sometimes entire. |
1–4(–8), axillary, 40–60 mm, 10–35(–40)-flowered, axis densely to sparsely eglandular-hairy, sometimes also glandular-hairy, rarely glabrate; bracts linear-lanceolate or narrowly ovate, (1.5–)3–4(–5) mm. |
Pedicels | erect-patent to spreading proximally, deflexed distally, (3–)6–9(–12) mm, shorter than subtending bract, densely glandular- and eglandular-hairy. |
erect, (0.5–)1–2(–3) mm, shorter than subtending bract, densely eglandular-hairy, rarely glandular-hairy or glabrate. |
Flowers | calyx lobes (5–)6–8.5(–10) mm, apex acuminate-acute, pairs connate 0.8–1.1 mm, glandular- and eglandular-hairy, rarely sparsely hairy; corolla mostly intense blue with white center, 4–6 mm diam.; stamens 2.5 mm; style (0.7–)1–1.3(–1.7) mm. |
calyx 4(or 5)-lobed, lobes 2–3.5(–4) mm, apex obtuse, glandular- and eglandular-hairy; corolla pale blue-lilac to pale pink, rarely white with pink veins, 6–8 mm diam.; stamens 2.8–3.4 mm; style 2.5–3(–4) mm. |
Capsules | slightly compressed in cross section, 3.5–5 × (4–)5–8 mm, apex emarginate, sinus angle 30–45(–60)°, densely eglandular- and glandular-hairy, rarely eglandular-hairy. |
compressed in cross section, triangulate-obdeltoid to obcordiform, 3–5 × 4–5 mm, apex rounded or truncate, rarely emarginate, glandular-hairy, rarely glabrous. |
Seeds | 4–10, yellowish brown, oblong to ovoid, cymbiform, 1.5–2.5 × 0.8–1.5 mm, 0.8 mm thick, cristate, dorsally reticulate-verrucate. |
9–24, brown to yellow, ellipsoid, ovoid, or obovoid, flat, 0.8–1.6 × (0.3–)0.6–1.2 mm, 0.2–0.3 mm thick, smooth. |
2n | = 18, 42 (Asia). |
= 18, 36 (Europe). |
Veronica argute-serrata |
Veronica officinalis |
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Phenology | Flowering Apr–Jun. | Flowering (Apr–)Jun–Jul(–Aug). |
Habitat | Bare soils, oak and juniper forests, steppes, alpine meadows, fields, stony slopes, streams, gardens, ruins, ruderal places, calcareous and siliceous soils. | Forests, meadows, pastures, scrub, ruderal places. |
Elevation | (600–)900–3000 m. ((2000–)3000–9800 ft.) | 0–1600 m. (0–5200 ft.) |
Distribution |
CO; ID; IL; KS; MT; NE; NM; NV; NY; OR; UT; WA; WY; BC; Eurasia [Introduced in North America]
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CA; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MT; NC; ND; NH; NJ; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; BC; NB; NF; NS; ON; PE; QC; SPM; Greenland; Eurasia [Introduced in North America; introduced in s South America (Chile)]
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Discussion | Veronica argute-serrata has been recorded mostly under the name V. biloba and occasionally under V. campylopoda Boissier. Most of the records for V. biloba likely belong under this name. Veronica biloba is also found in the flora area, sometimes mixed with V. argute-serrata; V. campylopoda has not been recorded from the flora area. A chromosome count of 2n = 18 (C. R. Bell 1965) for American material is suspicious because 2n = 42 has been reported multiple times in Turkish plants, and a base chromosome number of × = 9 is not known in related species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica ×tournefortii Villars [V. officinalis var. tournefortii (Villars) Dumortier] refers to the hybrid between V. officinalis and V. allionii Villars in Europe and is not applicable to any North American plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 321. | FNA vol. 17, p. 311. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | V. bornmuelleri | |
Name authority | Regel & Schmalhausen: Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 626. (1878) — (as argute serrata) | Linnaeus: Sp. Pl. 1: 11. (1753) |
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