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bilobed speedwell, sawtooth speedwell, twolobed speedwell

large-flower speedwell

Habit Annuals. Perennials.
Stems

erect, (7–)10–20(–30) cm, eglandular- and/or glandular-hairy.

prostrate to erect, unbranched, (3–)5–15(–20) cm, viscid-hairy.

Leaves

blade lanceolate or elliptic, (6–)12–28(–35) × 3–12(–20) mm, base cuneate, margins serrate, teeth 4–7(–12) per side, apex acute, surfaces sparsely glandular-hairy or glabrate.

blade elliptic-ovate, 20–45 × 15–30 mm, 1–3 times as long as wide, base cuneate, margins ± crenate-serrate, apex acute to rounded, abaxial surface with articulate hairs or glabrous, adaxial hirsute, or surfaces sparsely ciliate only.

Racemes

1–8, terminal, usually reduced and replaced by 2 pseudodichotomous racemes with additional axillary racemes, 80–150 mm, (5–)10–25(–35)-flowered, axis eglandular- and/or glandular-hairy;

bracts lanceolate, 9–12 mm, proximals with margins serrate, distals sometimes entire.

1–4, axillary, 50–150 mm, 3–8-flowered, axis viscid-hairy;

bracts oblong, 3–5 mm.

Pedicels

erect-patent to spreading proximally, deflexed distally, (3–)6–9(–12) mm, shorter than subtending bract, densely glandular- and eglandular-hairy.

suberect, 8–10 mm, longer than subtending bract, viscid-hairy.

Flowers

calyx lobes (5–)6–8.5(–10) mm, apex acuminate-acute, pairs connate 0.8–1.1 mm, glandular- and eglandular-hairy, rarely sparsely hairy;

corolla mostly intense blue with white center, 4–6 mm diam.;

stamens 2.5 mm;

style (0.7–)1–1.3(–1.7) mm.

calyx lobes 4 mm, apex obtuse, hairy;

corolla bright blue to violet-blue, 8–9 mm diam.;

stamens 7 mm;

style 7–9 mm.

Capsules

slightly compressed in cross section, 3.5–5 × (4–)5–8 mm, apex emarginate, sinus angle 30–45(–60)°, densely eglandular- and glandular-hairy, rarely eglandular-hairy.

compressed in cross section, obovoid, 9–11 × 7–8 mm, apex emarginate, sparsely hairy.

Seeds

4–10, yellowish brown, oblong to ovoid, cymbiform, 1.5–2.5 × 0.8–1.5 mm, 0.8 mm thick, cristate, dorsally reticulate-verrucate.

unknown.

2n

= 18, 42 (Asia).

= ca. 48–50 (Russia).

Veronica argute-serrata

Veronica grandiflora

Phenology Flowering Apr–Jun. Flowering Jul–Aug.
Habitat Bare soils, oak and juniper forests, steppes, alpine meadows, fields, stony slopes, streams, gardens, ruins, ruderal places, calcareous and siliceous soils. Meadows, between stones, slopes, alpine slopes.
Elevation (600–)900–3000 m. ((2000–)3000–9800 ft.) 0–600 m. (0–2000 ft.)
Distribution
from FNA
CO; ID; IL; KS; MT; NE; NM; NV; NY; OR; UT; WA; WY; BC; Eurasia [Introduced in North America]
[WildflowerSearch map]
from FNA
AK; Asia (Japan, Russia)
[BONAP county map]
Discussion

Veronica argute-serrata has been recorded mostly under the name V. biloba and occasionally under V. campylopoda Boissier. Most of the records for V. biloba likely belong under this name. Veronica biloba is also found in the flora area, sometimes mixed with V. argute-serrata; V. campylopoda has not been recorded from the flora area. A chromosome count of 2n = 18 (C. R. Bell 1965) for American material is suspicious because 2n = 42 has been reported multiple times in Turkish plants, and a base chromosome number of × = 9 is not known in related species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 321. FNA vol. 17, p. 311.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. bornmuelleri V. aphylla var. grandiflora, V. grandiflora var. minor
Name authority Regel & Schmalhausen: Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 626. (1878) — (as argute serrata) Gaertner: Novi Comment. Acad. Sci. Imp. Petrop. 14(1): 531, plate 18, fig. 1. (1770)
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