Veronica anagallis-aquatica |
Veronica persica |
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blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell |
bird's-eye speedwell, bird-eye speedwell, large field or bird's-eye speedwell, Persian speedwell, véronique de perse |
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Habit | Annuals or perennials. | Annuals. |
Stems | erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous. |
creeping to decumbent, 10–50(–60) cm, eglandular-hairy. |
Leaves | petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally; blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy. |
blade suborbiculate, broadly ovate, or broadly lanceolate, (6–)9–18(–30) × (5–)8–15(–20) mm, base truncate, margins serrate, apex acute, surfaces sparsely eglandular-hairy. |
Racemes | 6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy; bracts linear to lanceolate, 2–5 mm, apex acute. |
1–6, terminal, 100–500(–600) mm, 5–30-flowered, axis eglandular-hairy; bracts suborbiculate or broadly ovate or broadly lanceolate, (6–)9–18(–25) mm. |
Pedicels | suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy. |
spreading, deflexed in fruit, (12–)15–27(–38) mm, length 1–2(–3) times subtending bract, densely eglandular-hairy. |
Flowers | calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy; corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.; stamens 3–3.5 mm; style 1.5–2.5 mm. |
calyx lobes (4.5–)5.5–8(–9.5) mm, (1.7–)2.4–3.6(–4.2) mm wide, apex acuminate, eglandular-hairy; corolla intense bright blue, 8–14 mm diam.; stamens 1.2 mm; style (1.5–)2–2.8(–3.2) mm. |
Capsules | slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy. |
compressed in cross section, broadly obcordiform, 4–6 × (5–)6–8.5(–9.5) mm, apex acute, sinus angle (80–)90–120(–150)°, reticulate with prominent veins, ± sparsely to densely eglandular- and/or glandular-hairy or glabrate. |
Seeds | 40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
(10–)12–18(–20), pale brownish yellow, ellipsoid to globular, cymbiform, (1.3–)1.4–2.3(–2.5) × (0.8–)0.9–1.6(–1.9) mm, 0.5–1 mm thick, cristate-rugose. |
2n | = 36. |
= 28. |
Veronica anagallis-aquatica |
Veronica persica |
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Phenology | Flowering Mar–Sep(–Nov). | Flowering (Jan–)Apr–Jul(–Dec). |
Habitat | Stream margins, ditches, banks, springs, swamps, wet meadows. | Roadsides, lawns, fields, waste places. |
Elevation | 0–4000 m. (0–13100 ft.) | 0–500(–2000) m. (0–1600(–6600) ft.) |
Distribution |
AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
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AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; sw Asia [Introduced in North America; introduced also in Mexico (Michoacán, Veracruz), Central America, South America, Eurasia, e Asia, Africa, Atlantic Islands, Pacific Islands, Australia]
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Discussion | Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The names Veronica buxbaumii Tenore and V. tournefortii C. C. Gmelin (not V. ×tournefortii Villars) have been used for V. persica. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 313. | FNA vol. 17, p. 319. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Synonyms | V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera | V. persica var. aschersoniana, V. persica var. corrensiana, V. tournefortii subsp. aschersoniana, V. tournefortii subsp. corrensiana |
Name authority | Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) | Poiret: in J. Lamarck et al., Encycl. 8: 542. (1808) |
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