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blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell

tail kittentails

Habit Annuals or perennials.
Stems

erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous.

Leaves

petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally;

blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy.

persistent, some withering in 2d year as new leaves expand;

blade orbiculate to reniform, 25+ mm wide, not leathery, base cordate to lobate, margins ± incised-crenate to dentate, teeth apices rounded to acute or acuminate, surfaces glabrous, rarely sparsely hairy;

basal veins extending into distal 1/2 of blade, lateral veins 2–4 on each side of midvein.

Racemes

6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy;

bracts linear to lanceolate, 2–5 mm, apex acute.

erect, to 33 cm in fruit;

sterile bracts 3+, ovate-spatulate, largest 1+ cm;

flowers 12–100, loosely aggregated.

Pedicels

suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy.

Flowers

calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy;

corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.;

stamens 3–3.5 mm;

style 1.5–2.5 mm.

Stamens

epipetalous.

Ovaries

ovules 10–16.

Capsules

slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy.

glabrous or glabrescent.

Seeds

40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth.

Sepals

4.

Petals

(3 or)4(or 5), apex entire or erose;

corolla blue, ± regular, campanulate, much longer than calyx, glabrous or puberulent, tube conspicuous.

2n

= 36.

Veronica anagallis-aquatica

Synthyris missurica

Phenology Flowering Mar–Sep(–Nov).
Habitat Stream margins, ditches, banks, springs, swamps, wet meadows.
Elevation 0–4000 m. (0–13100 ft.)
Distribution
from FNA
AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; OR; WA
[WildflowerSearch map]
Discussion

Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Racemes usually 20–100-flowered, to 33 cm at end of flowering; ovaries glabrous; capsules glabrous.
S. missurica subsp. major
1. Racemes usually 12–45-flowered, to 25 cm at end of flowering; ovaries sparsely hairy, especially along margins; capsules glabrescent.
→ 2
2. Leaf blade margins regularly toothed; racemes to 20 cm at end of flowering.
S. missurica subsp. missurica
2. Leaf blade margins regularly to irregularly toothed; racemes to 25 cm at end of flowering.
S. missurica subsp. stellata
Source FNA vol. 17, p. 313. FNA vol. 17, p. 300.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Synthyris
Sibling taxa
V. agrestis, V. alpina, V. americana, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
S. alpina, S. borealis, S. bullii, S. canbyi, S. cordata, S. dissecta, S. laciniata, S. lanuginosa, S. missurica, S. oblongifolia, S. pinnatifida, S. plantaginea, S. platycarpa, S. ranunculina, S. reniformis, S. ritteriana, S. rubra, S. schizantha, S. wyomingensis
Subordinate taxa
S. missurica subsp. major, S. missurica subsp. missurica, S. missurica subsp. stellata
Synonyms V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera Veronica missurica
Name authority Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) (Rafinesque) Pennell: Proc. Acad. Nat. Sci. Philadelphia 85: 89. (1933)
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