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blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell

long-leaf speedwell, véronique à longues feuilles

Habit Annuals or perennials. Perennials.
Stems

erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous.

erect, usually with 10–20 nodes, (50–)70–120(–150) cm, proximal 1/2 glabrous, distal 1/2 eglandular-hairy with some glandular hairs intermixed.

Leaves

petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally;

blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy.

blade narrowly ovate-lanceolate, almost linear, or narrowly ovate, (60–)80–120(–150) × 10–30(–35) mm, base cuneate, truncate, or cordate, margins serrate to biserrate, apex acute, surfaces sparsely or densely eglandular-hairy, rarely glabrous.

Racemes

6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy;

bracts linear to lanceolate, 2–5 mm, apex acute.

1–7(–9), terminal and occasionally axillary, 100–200(–300) mm, 150–300-flowered, axis eglandular- and glandular-hairy;

bracts linear to subulate, 2–6 mm.

Pedicels

suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy.

erect, (1–)1.5–3(–5) mm, shorter than subtending bract, densely eglandular-hairy, sometimes also glandular-hairy.

Flowers

calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy;

corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.;

stamens 3–3.5 mm;

style 1.5–2.5 mm.

calyx lobes 2–2.5(–4) mm, apex acute, eglandular-hairy, sometimes glandular-hairy;

corolla blue, campanulate, longer than wide, 5–7 mm diam.;

stamens 5–6 mm;

style (5–)6–8(–10) mm.

Capsules

slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy.

± compressed in cross section, ovoid, 2.5–3 × 2.5–3 mm, apex emarginate, glabrous.

Seeds

40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth.

2–40, light brown, ellipsoid, flat, 0.7–1.2 × 0.5–0.8 mm, 0.2–0.3 mm thick, smooth.

2n

= 36.

= 34, 68 (Eurasia).

Veronica anagallis-aquatica

Veronica longifolia

Phenology Flowering Mar–Sep(–Nov). Flowering Jun–Aug(–Sep).
Habitat Stream margins, ditches, banks, springs, swamps, wet meadows. Moist (swampy) habitats, roadsides.
Elevation 0–4000 m. (0–13100 ft.) 0–800 m. (0–2600 ft.)
Distribution
from FNA
AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CT; FL; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; ND; NH; NJ; NY; OH; PA; RI; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Eurasia [Introduced in North America; introduced also in South America, Pacific Islands (New Zealand)]
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Reports of Veronica bachofenii Heuffel, V. grandis Fischer ex Sprengel (= V. daurica Steven), or V. spuria Linnaeus, seem always to refer to garden varieties of V. longifolia. Garden varieties often differ from typical plants in having glandular hairs and broader leaves, which may indicate past hybridization with other species.

Subgenus Pseudolysimachium includes Veronica longifolia and V. spicata, which are naturalized in the flora area, but also other species and hybrids of these, which are commonly grown ornamentally and may escape from cultivation. All of these plants should easily key out with V. longifolia and V. spicata but differ in specifics of the species descriptions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 313. FNA vol. 17, p. 308.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera V. maritima
Name authority Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) Linnaeus: Sp. Pl. 1: 10. (1753)
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