Veronica anagallis-aquatica |
Veronica dillenii |
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blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell |
Dillenius' speedwell |
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Habit | Annuals or perennials. | Annuals. |
Stems | erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous. |
erect to ascending, (8–)10–20(–40) cm, glandular- and eglandular-hairy. |
Leaves | petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally; blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy. |
blade lanceolate to ovate, 7–19(–21) × (3–)5–12 mm, 1–2 times as long as wide, base cuneate, margins (proximal) crenate-dentate or (distal) ±palmatifid, lobes 3–7+, apex of central lobe obtuse, lateral ± acute, surfaces glandular- and eglandular-hairy. |
Racemes | 6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy; bracts linear to lanceolate, 2–5 mm, apex acute. |
1, terminal, 60–180 mm, 15–50(–120)-flowered, axis eglandular- and glandular-hairy; proximal bracts similar to leaves, distal ones linear-lanceolate, 3–5 mm. |
Pedicels | suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy. |
erect, 2–4(–5) mm, shorter than subtending bract, length 1/2–1 times calyx, eglandular- and glandular-hairy. |
Flowers | calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy; corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.; stamens 3–3.5 mm; style 1.5–2.5 mm. |
calyx lobes 3.5–6 mm, apex acute, eglandular- and glandular-hairy; corolla deep blue, 4–6 mm diam.; stamens 0.8–1.5 mm; style 0.8–1.5 mm, stigma violet. |
Capsules | slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy. |
compressed in cross section, narrowly obcordiform, 3.5–4.5 × 4–6 mm, apex emarginate, angle of sinus ca. 90°, ciliate with glandular and eglandular hairs. |
Seeds | 40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
10–28, yellowish, ellipsoid, flat, 0.9–1.6 × 0.7–1.3 mm, 0.2–0.3 mm thick, smooth. |
2n | = 36. |
= 16 (Eurasia). |
Veronica anagallis-aquatica |
Veronica dillenii |
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Phenology | Flowering Mar–Sep(–Nov). | Flowering (Mar–)Apr–Jul. |
Habitat | Stream margins, ditches, banks, springs, swamps, wet meadows. | Open pine and oak forests, rocky, dry, sandy slopes. |
Elevation | 0–4000 m. (0–13100 ft.) | 500–2200 m. (1600–7200 ft.) |
Distribution |
AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
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IL; IN; MI; NY; VA; WI; Eurasia [Introduced in North America] |
Discussion | Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica dillenii is closely related to V. verna but with larger flowers; it may have been overlooked and may be distributed more widely. Most herbarium specimens of V. dillenii blacken when dry due to the presence of aucubin, which distinguishes them from V. verna. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 313. | FNA vol. 17, p. 318. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Synonyms | V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera | |
Name authority | Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) | Crantz: Stirp. Austr. Fasc. ed. 2, 2: 352. (1769) |
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