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blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell

chain speedwell, pink water speedwell, sessile water-speedwell, water speedwell

Habit Annuals or perennials. Annuals or perennials.
Stems

erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous.

erect or ascending, 15–60(–80) cm, glabrous or glandular-hairy distally.

Leaves

petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally;

blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy.

petiole 0 mm;

blade oblong-ovate to oblong-lanceolate, (5–)25–50(–100) × 4–15(–30) mm, 2.5–5 times as long as wide, base amplexical-truncate or amplexical-subcordate, margins entire or subentire, apex acute, surfaces glabrous.

Racemes

6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy;

bracts linear to lanceolate, 2–5 mm, apex acute.

10–25, axillary, 100–160 mm, 15–25-flowered, axis glabrous or glandular-hispid;

pedicels less than 3 per cm, glabrous or scarcely to densely short glandular-hairy, rarely completely glabrous;

bracts oblong, 3–5 mm, apex obtuse.

Pedicels

suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy.

patent, (3–)5–10 mm, equal to or ± shorter than subtending bract, glabrous or glandular-hairy.

Flowers

calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy;

corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.;

stamens 3–3.5 mm;

style 1.5–2.5 mm.

calyx lobes 2.5–3 mm, apex obtuse, glabrous or glandular-hairy;

corolla white to pale pink with darker veins not reaching margins, 4–5 mm diam.;

stamens 5 mm;

style (1.3–)1.5–2.5 mm.

Capsules

slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy.

± compressed in cross section, subglobular, 2.5–3(–3.5) × 3–4 mm, apex emarginate, sinus 0.1–0.3 mm, glabrous or glandular-hairy.

Seeds

40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth.

26–123, yellow-brown, ellipsoid to subglobular, planoconvex, 0.4–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth.

2n

= 36.

= 36.

Veronica anagallis-aquatica

Veronica catenata

Phenology Flowering Mar–Sep(–Nov). Flowering Jun–Oct.
Habitat Stream margins, ditches, banks, springs, swamps, wet meadows. Wet places, rarely running water, lakeshores, ditches, muddy places, stream channels.
Elevation 0–4000 m. (0–13100 ft.) 0–2500 m. (0–8200 ft.)
Distribution
from FNA
AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe
[WildflowerSearch map]
Discussion

Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica catenata seems to be a relative of V. anagallis-aquatica that is more cold-adapted, as seen in Europe (R. Götte 2007), and native to the flora area. Although no specimens were seen it may be found in Alaska, Connecticut, and Kentucky.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 313. FNA vol. 17, p. 314.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera V. catenata var. glandulosa, V. connata subsp. glaberrima
Name authority Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) Pennell: Rhodora 23: 37. (1921)
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