Veronica alpina |
Veronica anagallis-aquatica |
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alpine speedwell, véronique alpine |
blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell |
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Habit | Perennials. | Annuals or perennials. |
Stems | ascending, often dark bluish distally, sparsely branched at base, 5–15(–25) cm, eglandular-hairy. |
erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous. |
Leaves | blade ovate or oblong-elliptic, 7–30 × 4–20 mm, base cuneate, margins indistinctly dentate or entire, apex short-acuminate, abaxial surface glabrate, adaxial ± hairy. |
petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally; blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy. |
Racemes | 1, terminal, 5–20 mm, to 30 mm in fruit, (1–)3–20-flowered, axis eglandular-hairy; bracts oblanceolate, 4–5 mm. |
6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy; bracts linear to lanceolate, 2–5 mm, apex acute. |
Pedicels | ascending to erect, 1.5–2(–4) mm, shorter than subtending bract, eglandular-hairy. |
suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy. |
Flowers | calyx lobes 3–4 mm, ciliate, apex obtuse or acuminate, eglandular-hairy; corolla sky blue or blue-violet, sometimes white, rotate, 2.5–5.5 mm diam.; stamens 2–2.3 mm; style 0.5–1.5(–2) mm. |
calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy; corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.; stamens 3–3.5 mm; style 1.5–2.5 mm. |
Capsules | compressed in cross section, obovoid or oblong-obovoid, 4.5–7.5 × 3.5–5.5 mm, ca. as long as wide, apex shallowly emarginate, eglandular-hairy or glabrate. |
slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy. |
Seeds | 9–53, brown to yellow, ellipsoid-oblong, flat, 0.7–1.2 × 0.4–1 mm, 0.1 mm thick, smooth. |
40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
2n | = 18. |
= 36. |
Veronica alpina |
Veronica anagallis-aquatica |
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Phenology | Flowering (Jun–)Jul–Aug. | Flowering Mar–Sep(–Nov). |
Habitat | Slopes, moist rocks, hillocks, moist alpine and subalpine meadows. | Stream margins, ditches, banks, springs, swamps, wet meadows. |
Elevation | 0–1000 m. (0–3300 ft.) | 0–4000 m. (0–13100 ft.) |
Distribution |
NU; QC; Greenland; Eurasia |
AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
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Discussion | Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 17, p. 309. | FNA vol. 17, p. 313. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | V. alpina subsp. australis, V. alpina var. australis, V. pumila | V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera |
Name authority | Linnaeus: Sp. Pl. 1: 11. (1753) | Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) |
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