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field speedwell, green field speedwell, green procumbent speedwell, véronique agreste

alpine speedwell, American alpine speedwell, hairy speedwell, spring speedwell, thyme-leaf speedwell, wormskjold's alpine speedwell

Habit Annuals. Perennials.
Stems

creeping to decumbent, 5–25 cm, hairy.

erect or ascending, light grayish green distally, unbranched, (3–)8–50 cm, sparsely to densely villous-hirsute, sometimes also glandular-hairy.

Leaves

blade elliptic-ovate to oblong, (6–)8–16(–19) × (3–)4–10(–13) mm, base truncate, margins serrate, teeth 4–6(–8) per side, apex obtuse to acute, abaxial surface ± more densely hairy than adaxial.

blade elliptic to lanceolate or oblong-ovate, 8–40 × 5–20 mm, base cuneate, margins entire, dentate, or serrate, apex short-acuminate, surfaces sparsely to densely villous-hirsute or glabrous.

Racemes

1–5, terminal, 50–250 mm, 5–15-flowered, axis eglandular-hairy, sometimes glandular-hairy;

bracts elliptic-ovate to oblong, (6–)9–16(–19) mm.

1, terminal, 5–40 mm, to 60(–150) mm in fruit, (2–)5–25-flowered, axis densely villous-hirsute and ± glandular-hairy;

bracts linear to lanceolate, 1–8 mm.

Pedicels

deflexed, (5–)6–10(–15) mm, ± shorter than subtending bracts, densely eglandular- and, sometimes, glandular-hairy distally.

erect, 2–10(–15) mm, ca. equal to subtending bract, densely villous-hirsute or glandular-hairy.

Flowers

calyx lobes linear-lanceolate, (5–)6–7 mm, 2.2–2.6 mm wide, apex rounded, sparsely eglandular- or glandular-hairy;

corolla white or pale pinkish or pale blue, 4–5(–6) mm diam.;

stamens 0.5–1 mm;

style (0.6–)0.9–1.1(–1.2) mm.

calyx lobes (2.5–)3–5.5(–7) mm, apex obtuse or acuminate, glabrous or densely glandular-hairy;

corolla deep blue-violet or violet-purple to deep blue, rotate, 3–11 mm diam.;

stamens 1–2 mm;

style 0.8–4(–6) mm.

Capsules

± compressed in cross section, ± subglobular, 3.5–4.5(–4.7) × 4.5–6(–6.3) mm, apex emarginate, sparsely glandular-hairy.

compressed in cross section, oblong-obovoid, 4–6(–8) × (2.8–)4–5.5 mm, ca. as long as wide, apex emarginate, usually densely glandular-hairy.

Seeds

6–14, yellow to ochre, globular, cymbiform, 1.3–2.1 × 1–1.6 mm, 0.6–1.1 mm thick, strongly rugose (transversely ribbed).

10–50, straw colored, ellipsoid, flat, 0.7–1 × 0.4–1 mm, 0.05–1 mm thick, very minutely striate.

2n

= 28 (Europe).

Veronica agrestis

Veronica wormskjoldii

Phenology Flowering May–Oct.
Habitat Light, noncalcareous, moist soils, gravelly soils.
Elevation (0–)300–800 m. ((0–)1000–2600 ft.)
Distribution
from FNA
DC; FL; IL; LA; MA; NY; PA; TX; VT; NF; NS; QC; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; ME; MT; NH; NM; NV; OR; UT; WA; WY; AB; BC; NT; NU; ON; QC; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Veronica agrestis is similar to the more frequent V. persica and probably frequently overlooked and to be expected elsewhere. However, it should be noted that it is less common than V. persica or V. polita and always in more humid habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Veronica wormskjoldii occurs in two cytotypes, a diploid (2n = 18) in western North America and a tetraploid (2n = 36) in eastern North America and Greenland. As demonstrated by D. C. Albach et al. (2006), the tetraploid is likely to be a hybrid of the diploid V. alpina and V. wormskjoldii. In that publication, species rank was used for simplicity, not as a taxonomic conclusion. However, subsequent morphological analyses in conjunction with preparing this treatment revealed that the large variation in the diploid V. wormskjoldii, also supported by the large number of varieties established within that taxon, make it nearly impossible to differentiate the two North American taxa morphologically. In cases where two taxa are morphologically undifferentiable, but karyologically and genetically distinct and geographically clearly separate, the rank of subspecies seems more appropriate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pedicels 2–5 mm; e North America.
subsp. wormskjoldii
1. Pedicels (2–)5–10(–15) mm; w North America.
subsp. nutans
Source FNA vol. 17, p. 320. FNA vol. 17, p. 309.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna
Subordinate taxa
V. wormskjoldii subsp. nutans, V. wormskjoldii subsp. wormskjoldii
Synonyms V. alpina subsp. wormskjoldii, V. alpina var. wormskjoldii
Name authority Linnaeus: Sp. Pl. 1: 13. (1753) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 1: 101. (1817) — (as wormskjoldi)
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