Veronica agrestis |
Veronica wormskjoldii |
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field speedwell, green field speedwell, green procumbent speedwell, véronique agreste |
alpine speedwell, American alpine speedwell, hairy speedwell, spring speedwell, thyme-leaf speedwell, wormskjold's alpine speedwell |
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Habit | Annuals. | Perennials. | ||||
Stems | creeping to decumbent, 5–25 cm, hairy. |
erect or ascending, light grayish green distally, unbranched, (3–)8–50 cm, sparsely to densely villous-hirsute, sometimes also glandular-hairy. |
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Leaves | blade elliptic-ovate to oblong, (6–)8–16(–19) × (3–)4–10(–13) mm, base truncate, margins serrate, teeth 4–6(–8) per side, apex obtuse to acute, abaxial surface ± more densely hairy than adaxial. |
blade elliptic to lanceolate or oblong-ovate, 8–40 × 5–20 mm, base cuneate, margins entire, dentate, or serrate, apex short-acuminate, surfaces sparsely to densely villous-hirsute or glabrous. |
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Racemes | 1–5, terminal, 50–250 mm, 5–15-flowered, axis eglandular-hairy, sometimes glandular-hairy; bracts elliptic-ovate to oblong, (6–)9–16(–19) mm. |
1, terminal, 5–40 mm, to 60(–150) mm in fruit, (2–)5–25-flowered, axis densely villous-hirsute and ± glandular-hairy; bracts linear to lanceolate, 1–8 mm. |
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Pedicels | deflexed, (5–)6–10(–15) mm, ± shorter than subtending bracts, densely eglandular- and, sometimes, glandular-hairy distally. |
erect, 2–10(–15) mm, ca. equal to subtending bract, densely villous-hirsute or glandular-hairy. |
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Flowers | calyx lobes linear-lanceolate, (5–)6–7 mm, 2.2–2.6 mm wide, apex rounded, sparsely eglandular- or glandular-hairy; corolla white or pale pinkish or pale blue, 4–5(–6) mm diam.; stamens 0.5–1 mm; style (0.6–)0.9–1.1(–1.2) mm. |
calyx lobes (2.5–)3–5.5(–7) mm, apex obtuse or acuminate, glabrous or densely glandular-hairy; corolla deep blue-violet or violet-purple to deep blue, rotate, 3–11 mm diam.; stamens 1–2 mm; style 0.8–4(–6) mm. |
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Capsules | ± compressed in cross section, ± subglobular, 3.5–4.5(–4.7) × 4.5–6(–6.3) mm, apex emarginate, sparsely glandular-hairy. |
compressed in cross section, oblong-obovoid, 4–6(–8) × (2.8–)4–5.5 mm, ca. as long as wide, apex emarginate, usually densely glandular-hairy. |
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Seeds | 6–14, yellow to ochre, globular, cymbiform, 1.3–2.1 × 1–1.6 mm, 0.6–1.1 mm thick, strongly rugose (transversely ribbed). |
10–50, straw colored, ellipsoid, flat, 0.7–1 × 0.4–1 mm, 0.05–1 mm thick, very minutely striate. |
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2n | = 28 (Europe). |
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Veronica agrestis |
Veronica wormskjoldii |
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Phenology | Flowering May–Oct. | |||||
Habitat | Light, noncalcareous, moist soils, gravelly soils. | |||||
Elevation | (0–)300–800 m. ((0–)1000–2600 ft.) | |||||
Distribution |
DC; FL; IL; LA; MA; NY; PA; TX; VT; NF; NS; QC; Europe [Introduced in North America]
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AK; AZ; CA; CO; ID; ME; MT; NH; NM; NV; OR; UT; WA; WY; AB; BC; NT; NU; ON; QC; YT
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Discussion | Veronica agrestis is similar to the more frequent V. persica and probably frequently overlooked and to be expected elsewhere. However, it should be noted that it is less common than V. persica or V. polita and always in more humid habitats. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Veronica wormskjoldii occurs in two cytotypes, a diploid (2n = 18) in western North America and a tetraploid (2n = 36) in eastern North America and Greenland. As demonstrated by D. C. Albach et al. (2006), the tetraploid is likely to be a hybrid of the diploid V. alpina and V. wormskjoldii. In that publication, species rank was used for simplicity, not as a taxonomic conclusion. However, subsequent morphological analyses in conjunction with preparing this treatment revealed that the large variation in the diploid V. wormskjoldii, also supported by the large number of varieties established within that taxon, make it nearly impossible to differentiate the two North American taxa morphologically. In cases where two taxa are morphologically undifferentiable, but karyologically and genetically distinct and geographically clearly separate, the rank of subspecies seems more appropriate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 320. | FNA vol. 17, p. 309. | ||||
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. alpina subsp. wormskjoldii, V. alpina var. wormskjoldii | |||||
Name authority | Linnaeus: Sp. Pl. 1: 13. (1753) | Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 1: 101. (1817) — (as wormskjoldi) | ||||
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