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field speedwell, green field speedwell, green procumbent speedwell, véronique agreste

Copeland's speedwell

Habit Annuals. Perennials.
Stems

creeping to decumbent, 5–25 cm, hairy.

ascending, unbranched, 5–15 cm, densely glandular-hairy.

Leaves

blade elliptic-ovate to oblong, (6–)8–16(–19) × (3–)4–10(–13) mm, base truncate, margins serrate, teeth 4–6(–8) per side, apex obtuse to acute, abaxial surface ± more densely hairy than adaxial.

blade oblong-elliptic, (5–)10–15(–35) × 4–8 mm, base cuneate, margins entire, apex short-acuminate, surfaces hairy.

Racemes

1–5, terminal, 50–250 mm, 5–15-flowered, axis eglandular-hairy, sometimes glandular-hairy;

bracts elliptic-ovate to oblong, (6–)9–16(–19) mm.

1, terminal, distalmost leaves often with 1 or 2 axillary flowers, 10–80 mm, (3–)5–15-flowered, axis glandular-hairy;

bracts lanceolate, 3–5 mm.

Pedicels

deflexed, (5–)6–10(–15) mm, ± shorter than subtending bracts, densely eglandular- and, sometimes, glandular-hairy distally.

erect, 5–8 mm, equal to ± longer than subtending bract, densely glandular-hairy.

Flowers

calyx lobes linear-lanceolate, (5–)6–7 mm, 2.2–2.6 mm wide, apex rounded, sparsely eglandular- or glandular-hairy;

corolla white or pale pinkish or pale blue, 4–5(–6) mm diam.;

stamens 0.5–1 mm;

style (0.6–)0.9–1.1(–1.2) mm.

calyx 4(or 5)-lobed, lobes (1–)2–3 mm, apex obtuse, glandular-hairy;

corolla pale blue to purple, rotate, 8–10 mm diam.;

stamens 4–5 mm;

style 7 mm.

Capsules

± compressed in cross section, ± subglobular, 3.5–4.5(–4.7) × 4.5–6(–6.3) mm, apex emarginate, sparsely glandular-hairy.

compressed in cross section, broadly oblong, 5–6 × 3.5 mm, longer than wide, apex emarginate, glandular-hairy.

Seeds

6–14, yellow to ochre, globular, cymbiform, 1.3–2.1 × 1–1.6 mm, 0.6–1.1 mm thick, strongly rugose (transversely ribbed).

number unknown, brown, ovoid, flat, 1–1.2 × 0.7–1.1 mm, thickness and texture unknown.

2n

= 28 (Europe).

= 18.

Veronica agrestis

Veronica copelandii

Phenology Flowering May–Oct. Flowering Jul–Aug.
Habitat Light, noncalcareous, moist soils, gravelly soils. Subalpine meadows, alpine slopes.
Elevation (0–)300–800 m. ((0–)1000–2600 ft.) 1900–2500 m. (6200–8200 ft.)
Distribution
from FNA
DC; FL; IL; LA; MA; NY; PA; TX; VT; NF; NS; QC; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Veronica agrestis is similar to the more frequent V. persica and probably frequently overlooked and to be expected elsewhere. However, it should be noted that it is less common than V. persica or V. polita and always in more humid habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica copelandii is sister to V. cusickii in the phylogenetic analysis by D. C. Albach et al. (2006). It occurs in the Klamath Ranges of northwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 320. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Name authority Linnaeus: Sp. Pl. 1: 13. (1753) Eastwood: Bot. Gaz. 41: 288, fig. 2. (1906) — (as copelandi)
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