Veronica agrestis |
Veronica anagallis-aquatica |
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field speedwell, green field speedwell, green procumbent speedwell, véronique agreste |
blue speedwell, blue water speedwell, brook pimpernel, véronique mouron-d'eau, water pimpernel, water speedwell |
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Habit | Annuals. | Annuals or perennials. |
Stems | creeping to decumbent, 5–25 cm, hairy. |
erect or prostrate basally, (20–)30–100(–170) cm, usually thick-fleshy, glabrous. |
Leaves | blade elliptic-ovate to oblong, (6–)8–16(–19) × (3–)4–10(–13) mm, base truncate, margins serrate, teeth 4–6(–8) per side, apex obtuse to acute, abaxial surface ± more densely hairy than adaxial. |
petiole 0–2(–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally; blade (of proximal leaves) ovate, elliptic, or oblong, (15–)30–80(–145) × (7–)10–30(–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy. |
Racemes | 1–5, terminal, 50–250 mm, 5–15-flowered, axis eglandular-hairy, sometimes glandular-hairy; bracts elliptic-ovate to oblong, (6–)9–16(–19) mm. |
6–25, axillary, 50–100(–150) mm, 5–10 mm diam., (20–)30–40(–60)-flowered, axis glabrous, rarely glandular-hairy; bracts linear to lanceolate, 2–5 mm, apex acute. |
Pedicels | deflexed, (5–)6–10(–15) mm, ± shorter than subtending bracts, densely eglandular- and, sometimes, glandular-hairy distally. |
suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7(–10) mm, longer than subtending bracts, glabrous or hairy. |
Flowers | calyx lobes linear-lanceolate, (5–)6–7 mm, 2.2–2.6 mm wide, apex rounded, sparsely eglandular- or glandular-hairy; corolla white or pale pinkish or pale blue, 4–5(–6) mm diam.; stamens 0.5–1 mm; style (0.6–)0.9–1.1(–1.2) mm. |
calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy; corolla lavender to pale blue, rarely pale lilac, (4–)6–8 mm diam.; stamens 3–3.5 mm; style 1.5–2.5 mm. |
Capsules | ± compressed in cross section, ± subglobular, 3.5–4.5(–4.7) × 4.5–6(–6.3) mm, apex emarginate, sparsely glandular-hairy. |
slightly compressed in cross section, globular, (2.5–)3–3.5(–4) × 2.5–3.2(–4) mm, apex rounded or ± emarginate, glabrous or sparsely short glandular-hairy. |
Seeds | 6–14, yellow to ochre, globular, cymbiform, 1.3–2.1 × 1–1.6 mm, 0.6–1.1 mm thick, strongly rugose (transversely ribbed). |
40–77, yellow-brown, ellipsoid to subglobular, planoconvex, (0.3–)0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. |
2n | = 28 (Europe). |
= 36. |
Veronica agrestis |
Veronica anagallis-aquatica |
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Phenology | Flowering May–Oct. | Flowering Mar–Sep(–Nov). |
Habitat | Light, noncalcareous, moist soils, gravelly soils. | Stream margins, ditches, banks, springs, swamps, wet meadows. |
Elevation | (0–)300–800 m. ((0–)1000–2600 ft.) | 0–4000 m. (0–13100 ft.) |
Distribution |
DC; FL; IL; LA; MA; NY; PA; TX; VT; NF; NS; QC; Europe [Introduced in North America]
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AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; ME; MI; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; Eurasia; Africa; Atlantic Islands [Introduced in North America; likely introduced also in Mexico, Central America, South America]
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Discussion | Veronica agrestis is similar to the more frequent V. persica and probably frequently overlooked and to be expected elsewhere. However, it should be noted that it is less common than V. persica or V. polita and always in more humid habitats. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 320. | FNA vol. 17, p. 313. |
Parent taxa | Plantaginaceae > Veronica | Plantaginaceae > Veronica |
Sibling taxa | ||
Synonyms | V. anagallis-aquatica var. terrea, V. brittonii, V. comosa, V. glandifera, V. micromera | |
Name authority | Linnaeus: Sp. Pl. 1: 13. (1753) | Linnaeus: Sp. Pl. 1: 12. (1753) — (as anagall. s) |
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