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twiggy mullein, wand mullein

mullein

Habit Biennials. Herbs, annual, biennial, or perennial; stolons absent.
Stems

50–100 cm, densely stipitate-glandular, sometimes also sparsely hirsute-villous.

erect, glabrous, glabrate, puberulent, hirsute, tomentose, or floccose, sometimes glabrescent, stipitate-glandular or eglandular.

Leaves

surfaces densely stipitate-glandular, sometimes also sparsely hirsute-villous;

basal and proximal cauline with petiole 1–2 mm;

blade elliptic to elliptic-obovate, 8–20(–30) × 2.5–8(–15) cm, base subrounded to broadly cuneate;

cauline not clasping, gradually smaller distally, base not decurrent, margins coarsely crenate to crenulate, apex of distal cauline and floral bracts acute to obtuse.

basal and cauline, alternate;

stipules absent;

petiole present or absent;

blade not fleshy, leathery or not, margins entire, serrate, crenate, dentate, sinuate, or lobed.

Inflorescences

unbranched, narrowly cylindric, flowers remote, solitary in axils at least distally, sometimes 1(–5) at proximal nodes;

rachis densely stipitate-glandular, sometimes also sparsely hirsute-villous with simple hairs;

bracts linear-lanceolate, 8–20 mm, base not decurrent, apex long-acuminate, densely stipitate-glandular, sometimes also sparsely hirsute-villous with simple hairs.

terminal, spikes, racemes, or panicles, flowers solitary or clustered in fascicles;

bracts present.

Pedicels

free, (0–)1–3 mm;

bracteoles 2.

present;

bracteoles present or absent.

Flowers

calyx 4–9 mm, densely stipitate-glandular, sometimes also sparsely hirsute-villous with simple hairs, lobes ovate-lanceolate to triangular or narrowly lanceolate;

corolla yellow, (25–)30–40 mm diam., pellucid glands absent or relatively few;

proximal filaments glabrous at least distally, distal pair villous, hairs purplish to violet or whitish;

stigma capitate.

bisexual;

sepals 5, calyx radially or bilaterally symmetric, campanulate, lobes linear-oblong to elliptic or triangular;

petals 5, corolla yellow to orange, white, purple, cream, or pink, sometimes with purple center or red-tinged tips, radially or bilaterally symmetric, rotate with petals spreading to deflexed or shallowly cupulate, abaxial lobes 3, adaxial 2;

stamens [4]5, adnate to base of corolla, equal to subequal or anterior pair longer, filaments densely villous, sometimes anterior pair glabrous, staminode 0[1];

ovary 2-locular, placentation axile;

stigma capitate or spatulate.

Fruits

capsules, ovoid to ellipsoid-ovoid, broadly ellipsoid, ovoid-globular, or subglobular, dehiscence septicidal.

Capsules

ovoid-globular to subglobular, 6–10 mm, stipitate-glandular.

Seeds

50–300, tan or brown to orangish, conic to cylindric, usually pitted and rugose, appearing transversely and longitudinally ribbed, wings absent.

x

= 6.

Verbascum virgatum

Verbascum

Phenology Flowering Apr–Jun(–Oct).
Habitat Fields, roadsides, disturbed sites.
Elevation 10–2000 m. [0–6600 ft.]
Discussion

The occurrence of Verbascum virgatum in Nova Scotia may be historic; Ruth collected specimens from 1940 through 1960 from East Chester, Sydney, and Wolfville; it apparently has not been seen there subsequently. It also may be historic in British Columbia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 300–360 (12 in the flora).

Chromosome counts of 2n = 18, 26, 30, 32, 34, 36, 40, 44, 45, 48, 50, 64, and 66 have been reported for species of Verbascum. A base number is not clear, but x = 6 appears to be a reasonable fit (2n = 18, 30, 36, 48, and 66 would thus be polyploid levels). In any case, the range of numbers reflects both polyploidy and dysploidy.

Stamen number, placenta structure (entire and sessile versus two-fid and stalked), capsule shape, and number of flowers per node/bract have been used as diagnostic characters to distinguish Celsia Linnaeus and Staurophragma Fischer & C. A. Meyer as segregates from Verbascum (S. Murbeck 1925; A. Huber-Morath 1973; F. A. Karaveliogullari and Z. Aytac 2008). Huber-Morath (1978) included Celsia and Staurophragma within Verbascum. All species treated here are Verbascum in the strict sense.

Taxonomic treatments of Verbascum have been inconsistent in use of rank for infraspecific taxa; both varieties and subspecies appear in this treatment as well. Since all of our taxa are introduced to North America, further study of the species in their native environs is warranted before any uniformity can be established. In any case, use of one rank or the other does not imply a biological or evolutionary difference.

Hybrids are commonly produced among many combinations of parents. Verbascum thapsus, which is the most common and widespread of the species in the flora area, is known to have formed four hybrids in North America; see species discussions for nothospecies and purported parental species. Verbascum ×ramigerum Link ex Schrader [V. densiflorum × V. lychnitis] has been reported in Europe; it and other combinations may be expected in the flora area.

Plants of Verbascum collected as weeds in a lawn in Duluth, Minnesota, in 2001 and 2002 (Schimpf DJS318, DJS327, MIN), have been identified as V. chaixii Villars (http://plants.usda.gov); they perhaps are hybrids with V. nigrum as one of the parents (Dirk Albach, pers. comm.). The leaves are glabrate, the basal ones ovate-elliptic, subentire to shallowly crenate, and subpetiolate, and the cauline ones mostly non-clasping; the inflorescences are stipitate-glandular, unbranched, with flowers in loosely overlapping clusters of three to six; corollas are white to yellowish with pellucid glands; and staminal filaments all are villous with violet hairs. Without an unequivocal identification and evidence that they have become naturalized, they are not formally treated here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Distribution
from FNA
AL; AZ; CA; FL; GA; ID; IL; IN; LA; NC; NM; NV; NY; OH; PA; SC; TX; UT; BC; NS; ON; QC; Europe; Asia [Introduced in North America; introduced also in Mexico (Coahuila), South America (Argentina, Chile), s Asia (India), Pacific Islands (Hawaii, New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
from USDA
Europe; Asia; ne Africa [Introduced in North America; introduced also in Mexico, South America, elsewhere in Africa (Tunisia), Pacific Islands, Australia]
[BONAP county map]
Parent taxa Scrophulariaceae > Verbascum Scrophulariaceae
Sibling taxa
V. blattaria, V. bombyciferum, V. densiflorum, V. lychnitis, V. nigrum, V. phlomoides, V. phoeniceum, V. pulverulentum, V. sinuatum, V. speciosum, V. thapsus
Subordinate taxa
V. blattaria, V. bombyciferum, V. densiflorum, V. lychnitis, V. nigrum, V. phlomoides, V. phoeniceum, V. pulverulentum, V. sinuatum, V. speciosum, V. thapsus, V. virgatum
Key
1.Flowers solitary in axils at least distally.→ 2
2.Corollas purple to violet; bracteoles 0; cauline leaves abruptly smaller distally.V. phoeniceum
2.Corollas yellow, sometimes white or pink (V. blattaria); bracteoles 0 or 2; cauline leaves gradually smaller distally.→ 3
3.Pedicels 5–11(–15) mm; bracteoles 0; stems and leaves glabrous or glabrate.V. blattaria
3.Pedicels (0–)1–3 mm; bracteoles 2; stems and leaves densely stipitate-glandular, sometimes also sparsely hiruste-villous.V. virgatum
1.Flowers mostly in clusters of 2–10.→ 4
4.Inflorescences unbranched, sometimes branched from proximal nodes.→ 5
5.Basal and proximal cauline leaf blades: bases shallowly cordate to nearly truncate, surfaces sparsely tomentose to glabrate, abaxial soon glabrescent, sometimes both glabrate; filaments villous, hairs purple to violet.V. nigrum
5.Basal and proximal cauline leaf blades: bases attenuate, surfaces densely and persistently tomentose; filaments villous or glabrous, hairs yellowish to whitish.→ 6
6.Flower clusters loosely overlapping.V. bombyciferum
6.Flower clusters densely overlapping or remote proximally.→ 7
7.Cauline leaves: bases not decurrent, rarely slightly so.V. phlomoides
7.Cauline leaves: bases decurrent.→ 8
8.Corollas 14–20(–30) mm diam., pellucid glands relatively numerous; anthers yellow; stigmas capitate; pedicels mostly or completely adnate to rachis.V. thapsus
8.Corollas 30–55 mm diam., pellucid glands absent or relatively few; anthers orange; stigmas spatulate; pedicels free or adnate to rachis at base.V. densiflorum
4.Inflorescences freely branched, sometimes unbranched (V. densiflorum, V. nigrum) or branched from proximal nodes and forming panicles (V. nigrum).→ 9
9.Leaf surfaces persistently tomentose.→ 10
10.Filaments: proximals glabrous, distal pairs villous; cauline leaves: bases long-decurrent.V. densiflorum
10.Filaments villous; cauline leaves: bases not decurrent or short-decurrent.→ 11
11.Leaf blade margins sinuate to incised or incised-lobed; flowers remote, clustered, rarely solitary; pedicels 2–5 mm; filament hairs violet to purple.V. sinuatum
11.Leaf blade margins entire or minutely serrate-dentate; flowers loosely overlapping, clustered; pedicels (3–)5–12 mm; filament hairs whitish to yellowish.V. speciosum
9.Leaf surfaces glabrate, sometimes abaxial surfaces sparsely persistently hairy and soon glabrescent.→ 12
12.Basal and proximal cauline leaves shallowly cordate or nearly truncate to a narrow petiole 5–15(–20) mm; filament hairs purple to violet; inflorescences unbranched, sometimes branched from proximal nodes, narrowly conic panicles.V. nigrum
12.Basal leaves sessile or basally attenuate and sessile, less commonly with petiolar regions 10–50 mm, cauline leaves sessile; filament hairs yellow to whitish; inflorescences freely branched, loosely conic to broadly cylindric panicles, sometimes unbranched.→ 13
13.Cauline leaves: bases not clasping or slightly so, surfaces thinly tawny- to gray-tomentose, abaxial soon glabrescent, adaxial persistent, closely adherent; bracts linear to narrowly lanceolate, 8–15 mm; pedicels 6–11 mm; corollas white, sometimes yellowish.V. lychnitis
13.Cauline leaves: bases distinctly clasping or subclasping, surfaces densely and loosely white-floccose, glabrescent, especially abaxially, sometimes thin-persistent adaxially; bracts linear, 3–5 mm; pedicels (1–)2–5(–7) mm; corollas yellow.V. pulverulentum
Name authority Stokes: in W. Withering, Bot. Arr. Brit. Pl. ed. 2, 1: 227. (1787) Linnaeus: Sp. Pl. 1: 177. (1753): Gen. Pl. ed. 5, 83. (1754)
Source Flora of North America vol. 17, p. 347. Flora of North America vol. 17, p. 343.
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