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twiggy mullein, wand mullein

figwort family

Habit Biennials. Shrubs, subshrubs, trees, or herbs, annual, biennial, or perennial, not fleshy [fleshy], autotrophic.
Stems

50–100 cm, densely stipitate-glandular, sometimes also sparsely hirsute-villous.

prostrate, ascending, pendent, or erect.

Leaves

surfaces densely stipitate-glandular, sometimes also sparsely hirsute-villous;

basal and proximal cauline with petiole 1–2 mm;

blade elliptic to elliptic-obovate, 8–20(–30) × 2.5–8(–15) cm, base subrounded to broadly cuneate;

cauline not clasping, gradually smaller distally, base not decurrent, margins coarsely crenate to crenulate, apex of distal cauline and floral bracts acute to obtuse.

deciduous, semipersistent, or persistent, basal and cauline or cauline, opposite or alternate, simple;

stipules absent or present (most Buddleja, Emorya, Limosella);

petiole present or absent;

blade fleshy or not, leathery or not, margins entire to subentire, undulate, toothed, lobed, divided, or incised.

Inflorescences

unbranched, narrowly cylindric, flowers remote, solitary in axils at least distally, sometimes 1(–5) at proximal nodes;

rachis densely stipitate-glandular, sometimes also sparsely hirsute-villous with simple hairs;

bracts linear-lanceolate, 8–20 mm, base not decurrent, apex long-acuminate, densely stipitate-glandular, sometimes also sparsely hirsute-villous with simple hairs.

terminal, subterminal, or axillary, racemes, cymes, panicles, or thyrses (or combinations thereof), spikes, fascicles, or flowers 1(or 2).

Pedicels

free, (0–)1–3 mm;

bracteoles 2.

Flowers

calyx 4–9 mm, densely stipitate-glandular, sometimes also sparsely hirsute-villous with simple hairs, lobes ovate-lanceolate to triangular or narrowly lanceolate;

corolla yellow, (25–)30–40 mm diam., pellucid glands absent or relatively few;

proximal filaments glabrous at least distally, distal pair villous, hairs purplish to violet or whitish;

stigma capitate.

bisexual or unisexual (some Buddleja), perianth and androecium hypogynous;

sepals 4 or 5, ± distinct (Capraria), connate proximally, or to past middle (Limosella), calyx radially or bilaterally symmetric;

petals 4 or 5, proximally connate, corolla radially or bilaterally symmetric, regular or bilabiate, rotate to salverform, tubular, funnelform, or campanulate;

stamens mostly 4 or 5(–8 in Myoporum), adnate to corolla, didynamous or equal, staminode 0 or 1;

pistil 1, 2-carpellate, ovary superior, 2- or 4-locular (partition incomplete and ovary 1-locular distally in Limosella), placentation axile (free-central in Limosella, apical in Myoporum);

ovules anatropous or hemitropous (Buddleja), unitegmic, tenuinucellate;

style 1;

stigma 1, sometimes 2-lobed.

Fruits

capsules, dry and dehiscence septicidal or loculicidal, or fleshy and drupelike (Bontia, Myoporum) or berries (some Buddleja), [schizocarps].

Capsules

ovoid-globular to subglobular, 6–10 mm, stipitate-glandular.

Seeds

1–300, white, yellow, orangish, brown, or black, ovoid, oblong-ovoid, conic, ellipsoid, L-shaped, angled, cylindric, threadlike, or fusiform;

embryo straight or slightly curved, endosperm abundant or not.

Verbascum virgatum

Scrophulariaceae

Phenology Flowering Apr–Jun(–Oct).
Habitat Fields, roadsides, disturbed sites.
Elevation 10–2000 m. [0–6600 ft.]
Discussion

The occurrence of Verbascum virgatum in Nova Scotia may be historic; Ruth collected specimens from 1940 through 1960 from East Chester, Sydney, and Wolfville; it apparently has not been seen there subsequently. It also may be historic in British Columbia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera ca. 60, species ca. 1700 (9 genera, 45 species in the flora).

As noted in the introduction to this volume, the authors follow a narrow circumscription of Scrophulariaceae. Molecular studies (especially R. G. Olmstead et al. 2001 and B. Oxelman et al. 2005) have shown that some of the taxa commonly included in earlier treatments of Scrophulariaceae are better placed elsewhere, particularly in Orobanchaceae (hemiparasitic members, for example, Rhinantheae Lamarck & de Candolle) and Plantaginaceae (for example, Penstemon). Oxelman et al. and D. C. Tank et al. (2006) both recognized eight tribes in their analyses of the Scrophulariaceae. Five of the eight, Buddlejeae Bartling (Buddleja, Emorya), Leucophylleae Miers (Capraria, Leucophyllum), Limoselleae Dumortier (Limosella), Myoporeae Reichenbach (Bontia, Myoporum), and Scrophularieae Dumortier (Scrophularia, Verbascum), are represented in the flora area.

Inclusion of Myoporaceae (three or four genera and 125 species, in the sense of A. Cronquist 1981) is warranted based on its close similarity to Leucophylleae. Morphological similarities have led workers either to propose transferring Leucophylleae from Scrophulariaceae to Myoporaceae (C. J. Niezgoda and A. S. Tomb 1975) or to question the validity of recognizing Myoporaceae as distinct from Scrophulariaceae (J. Henrickson and L. D. Flyr 1985). R. G. Olmstead et al. (2001) confirmed this closeness in their molecular survey, with Leucophyllum and Myoporum clustering together and forming a clade sister to Buddlejaceae; Leucophylleae and Myoporeae also clustered together in the studies by B. Oxelman et al. (2005) and E. Gándara and V. Sosa (2013).

Members of Buddlejaceae (10 genera, ca. 150 species, in the sense of A. Cronquist 1981) often have been included in Loganiaceae (G. K. Rogers 1986) or treated as a separate family. Cronquist noted that the four-lobed corolla may be the primitive condition for the Scrophulariales and that Buddlejaceae is clearly not primitive. B. Oxelman et al. (1999) showed that Buddlejaceae is monophyletic; R. G. Olmstead et al. (2001) and Oxelman et al. (2005) both found that Buddleja clustered within Scrophulariaceae in the strict sense; that interpretation is followed here. J. H. Chau et al. (2017) found that Buddlejeae is monophyletic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Distribution
from FNA
AL; AZ; CA; FL; GA; ID; IL; IN; LA; NC; NM; NV; NY; OH; PA; SC; TX; UT; BC; NS; ON; QC; Europe; Asia [Introduced in North America; introduced also in Mexico (Coahuila), South America (Argentina, Chile), s Asia (India), Pacific Islands (Hawaii, New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
nearly worldwide except boreal and arctic North America and Asia; tropical Africa; Antarctica
[BONAP county map]
Parent taxa Scrophulariaceae > Verbascum
Sibling taxa
V. blattaria, V. bombyciferum, V. densiflorum, V. lychnitis, V. nigrum, V. phlomoides, V. phoeniceum, V. pulverulentum, V. sinuatum, V. speciosum, V. thapsus
Subordinate taxa
Bontia, Buddleja, Capraria, Emorya, Leucophyllum, Limosella, Myoporum, Scrophularia, Verbascum
Key
1.Herbs or subshrubs.→ 2
2.Plants aquatic or paludal; stems prostrate or ascending; leaves basal; stolons present.Limosella
2.Plants terrestrial; stems erect; leaves basal and cauline or cauline; stolons absent.→ 3
3.Leaves opposite.Scrophularia
3.Leaves alternate.→ 4
4.Leaf surfaces with punctate glands and internal secretory oil cavities; inflorescences axillary, racemes.Capraria
4.Leaf surfaces without punctate glands and internal secretory oil cavities; inflorescences terminal, spikes, racemes, or panicles.Verbascum
1.Shrubs or trees.→ 5
5.Sepals 4; stipules usually present or as stipular lines.→ 6
6.Calyx lobes linear-subulate; corollas long-tubular.Emorya
6.Calyx lobes ovate to lanceolate; corollas campanulate-rotate, salverform, funnelform, or tubular.Buddleja
5.Sepals 5; stipules absent.→ 7
7.Stems densely silvery gray-tomentose, sometimes appressed-stellate; corollas purplish lavender to violet or blue, rarely white, with white in proximal throat, usually marked with yellow-brown, orange, or purple-violet spots; fruits capsules, dry.Leucophyllum
7.Stems glabrous; corollas white, spotted with purple, or buff or coppery brown to greenish tan; fruits drupelike capsules, fleshy.→ 8
8.Corollas white, spotted with purple.Myoporum
8.Corollas buff or coppery brown to greenish tan.Bontia
Name authority Stokes: in W. Withering, Bot. Arr. Brit. Pl. ed. 2, 1: 227. (1787) Jussieu
Source Flora of North America vol. 17, p. 347. Flora of North America vol. 17, p. 324.
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