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golden inside-out flower, Siskiyou inside-out-flower, yellow vancouveria

northern inside-out flower, northern vancouveria, white inside-out-flower

Leaves

persistent, 2-ternately compound or 3-5-foliolate, 10-18 cm;

petiole 3-12 cm, sparsely hairy.

falling when fruits maturing, 2-3-ternately compound, 8-30 cm;

petiole 3-25 cm, pilose at base.

Leaflet

blades ovate to oblong, slightly 3-lobed, leathery, base cordate, margins thickened, crisped, apex notched;

surfaces abaxially pubescent, glaucous, adaxially glabrous to rarely pubescent.

blades narrowly to broadly ovate to rhomboid or rounded pentagonal, often 3-lobed, base cordate, margins entire to slightly sinuate and not conspicuously thickened, apex rounded to notched;

surfaces abaxially sparsely hairy, adaxially glabrous.

Inflorescences

peduncle 2-3 dm;

pedicel 1-4 cm, stipitate-glandular.

peduncle 2-3 dm;

pedicel 1-3 cm, glands absent.

Flowers

4-15;

bracteoles 6-9, tan to brown, 1-4 mm, caducous, stipitate-glandular;

sepals 6, yellow, spatulate, 6-10 mm, stipitate-glandular;

petals 6, yellow, 4-6 mm, margins entire, apex strongly reflexed, apical nectary darker yellow;

filaments stipitate-glandular.

5-30;

bracteoles 6-9, white, yellowish when dried, dotted with glandular trichomes;

sepals 6, white, 5-12 mm;

petals 6, white, yellowish when dried, 4-6 mm, margins entire, petal apex strongly reflexed, with nectar-bearing pocket, nectaries golden;

filaments stipitate-glandular.

Seeds

3-10, reddish brown, reniform to oblong, 3-4 mm.

1-6, black, lunate to reniform, 3 mm.

Follicles

greenish brown, 8-15 mm including beak, beak 3-4 mm, densely stipitate-glandular.

greenish to light brown, 10-15 mm including beak, beak 2-3 mm, stipitate-glandular.

2n

= 12.

= 12.

Vancouveria chrysantha

Vancouveria hexandra

Phenology Flowering spring (May–Jun); fruiting spring–summer (Jun–Jul). Flowering and fruiting spring–summer (May–Jul).
Habitat Open, mixed evergreen forests and thickets on serpentine substrates Redwood and Douglas-fir forests, deep shade
Elevation 100-1500 m (300-4900 ft) 100-1700 m (300-5600 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 3. FNA vol. 3.
Parent taxa Berberidaceae > Vancouveria Berberidaceae > Vancouveria
Sibling taxa
V. hexandra, V. planipetala
V. chrysantha, V. planipetala
Synonyms Epimedium hexandrum
Name authority Greene: Bull. Calif. Acad. Sci. 1: 66. (1885) (Hooker) C. Morren & Decaisne: Ann. Sci. Nat., Bot., sér. 2, 2: 351. (1833)
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