Vancouveria chrysantha |
Vancouveria |
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golden inside-out flower, Siskiyou inside-out-flower, yellow vancouveria |
inside-out-flower |
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Habit | Herbs, perennial, evergreen or deciduous, 1-5 dm, glabrous, glandular-pubescent, or sparsely hairy. | |||||||||
Rhizomes | extensive, creeping, nodose, producing 3 or more foliage leaves and flowering shoots per year. |
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Aerial stems | absent. |
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Leaves | persistent, 2-ternately compound or 3-5-foliolate, 10-18 cm; petiole 3-12 cm, sparsely hairy. |
blade deltate in overall outline; rachis without pulvinae; leaflet blades rhomboid or rounded pentagonal to ovate to oblong, shallowly 3-lobed, margins entire to sinuate; venation palmate. |
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Leaflet | blades ovate to oblong, slightly 3-lobed, leathery, base cordate, margins thickened, crisped, apex notched; surfaces abaxially pubescent, glaucous, adaxially glabrous to rarely pubescent. |
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Inflorescences | peduncle 2-3 dm; pedicel 1-4 cm, stipitate-glandular. |
terminal, racemes or panicles, open. |
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Flowers | 4-15; bracteoles 6-9, tan to brown, 1-4 mm, caducous, stipitate-glandular; sepals 6, yellow, spatulate, 6-10 mm, stipitate-glandular; petals 6, yellow, 4-6 mm, margins entire, apex strongly reflexed, apical nectary darker yellow; filaments stipitate-glandular. |
3-merous, 6-14 mm; bracteoles 6-9, sepaloid; sepals 6, white to yellow; petals 6, white to yellow, hooded with tip reflexed or flat, bearing nectar; stamens 6; anthers dehiscing by 2 apically hinged flaps; pollen exine striate; ovaries ellipsoid; placentation marginal; style lateral. |
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Fruits | follicles, brown, asymmetric, generally elliptic, dehiscing by 2 valves. |
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Seeds | 3-10, reddish brown, reniform to oblong, 3-4 mm. |
4-7, black to reddish brown; aril whitish, covering ca. 1/2-2/3 of seed. |
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Follicles | greenish brown, 8-15 mm including beak, beak 3-4 mm, densely stipitate-glandular. |
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x | = 6. |
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2n | = 12. |
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Vancouveria chrysantha |
Vancouveria |
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Phenology | Flowering spring (May–Jun); fruiting spring–summer (Jun–Jul). | |||||||||
Habitat | Open, mixed evergreen forests and thickets on serpentine substrates | |||||||||
Elevation | 100-1500 m (300-4900 ft) | |||||||||
Distribution |
CA; OR
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w United States |
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Discussion | Species 3 (3 in the flora). The fruits of Vancouveria are thin-walled follicles that are green or greenish brown at the time of dehiscence. The follicles dehisce by means of two valves that begin below the style and open to the base. The two valves recurve, exposing the seeds downward. In V. hexandra the follicle opens before the seeds are mature. The green seeds continue to grow and ripen in the open follicle. The appendage or aril on Vancouveria seeds has been shown to elaiosome (R. Y. Berg 1972). Ants carry the seeds to their nests and harvest the appendage as a food source. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 3. | FNA vol. 3. | ||||||||
Parent taxa | Berberidaceae > Vancouveria | Berberidaceae | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | Greene: Bull. Calif. Acad. Sci. 1: 66. (1885) | C. Morren & Decaisne: Ann. Sci. Nat., Bot., sér. 2, 2: 351. (1834) | ||||||||
Web links |