Vancouveria chrysantha |
Berberidaceae |
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golden inside-out flower, Siskiyou inside-out-flower, yellow vancouveria |
barberry family |
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Habit | Herbs or shrubs [trees], perennial, evergreen or deciduous, sometimes rhizomatous. | |||||||||||||||||||||||||||||
Stems | with or without spines. |
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Leaves | persistent, 2-ternately compound or 3-5-foliolate, 10-18 cm; petiole 3-12 cm, sparsely hairy. |
alternate, opposite, or fascicled, simple, 2-3-foliolate, or 1-3-pinnately or 2-3(-4)-ternately compound; stipules present or absent; venation pinnate or palmate. |
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Leaflet | blades ovate to oblong, slightly 3-lobed, leathery, base cordate, margins thickened, crisped, apex notched; surfaces abaxially pubescent, glaucous, adaxially glabrous to rarely pubescent. |
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Inflorescences | peduncle 2-3 dm; pedicel 1-4 cm, stipitate-glandular. |
terminal or axillary, racemes, cymes, umbels (or umbel-like), spikes, or panicles, or flowers solitary or in pairs, flowers pedicellate or sessile. |
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Flowers | 4-15; bracteoles 6-9, tan to brown, 1-4 mm, caducous, stipitate-glandular; sepals 6, yellow, spatulate, 6-10 mm, stipitate-glandular; petals 6, yellow, 4-6 mm, margins entire, apex strongly reflexed, apical nectary darker yellow; filaments stipitate-glandular. |
bisexual, inconspicuous or showy, radially symmetric; stipitate glands absent (except in Vancouveria); sepaloid bracteoles 0-9; perianth sometimes absent (Achlys), more frequently present, 2- or 3-merous, or sepals and petals intergrading (Nandis); sepals 6, distinct, often petaloid and colored, not spurred; petals 6-9, distinct, plane or hooded; nectary present; stamens 6; anthers dehiscing by valves or longitudinal slits; ovary superior, apparently 1-carpellate; placentation marginal or appearing basal; style present or absent, sometimes persistent in fruit as beak. |
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Fruits | follicles, berries, or utricles. |
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Seeds | 3-10, reddish brown, reniform to oblong, 3-4 mm. |
1-50, sometimes arillate; endosperm abundant; embryo large or small; mature seeds elevated on elongating stalk in Caulophyllum. |
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Follicles | greenish brown, 8-15 mm including beak, beak 3-4 mm, densely stipitate-glandular. |
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2n | = 12. |
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Vancouveria chrysantha |
Berberidaceae |
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Phenology | Flowering spring (May–Jun); fruiting spring–summer (Jun–Jul). | |||||||||||||||||||||||||||||
Habitat | Open, mixed evergreen forests and thickets on serpentine substrates | |||||||||||||||||||||||||||||
Elevation | 100-1500 m (300-4900 ft) | |||||||||||||||||||||||||||||
Distribution |
CA; OR
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Widespread; well represented in the north temperate zone |
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Discussion | Genera 15, species ca. 650 (8 genera, 33 species in the flora). Berberidaceae presents several interesting biogeographic features. Achlys is disjunct from western North America to east Asia with few morphologic differences between taxa. Diphylleia, Jeffersonia, and Podophyllum, each with a single eastern North American species, exhibit wide disjunctions to east Asia. Caulophyllum has three species, one in east Asia and two in the flora. Vancouveria is endemic to northwestern United States with nearest relations to Epimedium Linnaeus (H. Loconte and J. R. Estes 1989b; W. T. Stearn 1938), an exclusively Eastern Hemisphere genus. Nandina, Berberis, Epimedium, and Podophyllum are cultivated. The perianth of Berberidaceae is commonly composed of three distinct types of organs, but terminology for the organs varies from author to author. In our treatment, we refer to the small, outer parts as bracteoles (collectively forming a calyculus); the large, middle parts as sepals; and the innermost parts, which are commonly nectariferous, as petals. Some authors have referred to the bracteoles as outer sepals and to the petals as staminodes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 3. | FNA vol. 3, p. 272. | ||||||||||||||||||||||||||||
Parent taxa | Berberidaceae > Vancouveria | |||||||||||||||||||||||||||||
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Name authority | Greene: Bull. Calif. Acad. Sci. 1: 66. (1885) | Jussieu | ||||||||||||||||||||||||||||
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