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para grass

paragrass, signalgrass

Habit Plants perennial; stoloniferous, straggling. Plants annual or perennial; usually cespitose, sometimes mat-forming, sometimes stoloniferous.
Culms

to 5 m long, long-decumbent and rooting at the lower nodes, vertical portion 90-200(300) cm;

nodes villous.

5-500 cm, herbaceous, erect, geniculate, or decumbent and rooting at the lower nodes.

Sheaths

open;

auricles rarely present;

ligules apparently of hairs, the basal membranous portion inconspicuous;

blades ovate-lanceolate to lanceolate, flat.

Panicles

10-25 cm long, 5-10 cm wide, pyramidal, with 10-30 spikelike branches in more than 2 ranks;

primary branches 2.5-8 cm long, 0.4-0.9 mm wide, ascending to divergent, axils pubescent, axes flat, glabrous or with a few papillose-based hairs, secondary branches present or absent;

pedicels shorter than the spikelets, scabrous, sometimes with hairs.

Inflorescences

terminal or terminal and axillary, usually panicles of spikelike primary branches in 2 or more ranks, rachises not concealed by the spikelets;

primary branches usually alternate or subopposite, spikelike, and 1-sided, less frequently verticillate, axes flat or triquetrous, usually terminating in a well-developed, rudimentary spikelet;

secondary branches present or absent, axes flat or triquetrous;

disarticulation beneath the spikelets.

Spikelets

2.6-3.5 mm long, 1-1.4 mm wide, mostly in pairs, in 2-4 rows, appressed to the branches, purplish to green.

solitary, paired, or in triplets, subsessile or pedicellate, divergent or appressed, ovoid to ellipsoid, dorsally compressed, in 1-2(4) rows, with 2 florets, lower or upper glumes adjacent to the branch axes.

Glumes

scarcely separate, rachilla internodes short not pronounced;

lower glumes 0.6-1.1 mm, 1/5 – 1/3 as long as the spikelets, glabrous, 0-1(3)-veined;

upper glumes 2.6-3.5 mm, glabrous, 5-(7)-veined, without cross venation;

lower florets staminate;

lower lemmas 2.6-3.3 mm, glabrous, 5-veined, without cross venation;

upper lemmas 2.3-2.8 mm long, 1-1.3 mm wide, apices rounded, mucronate;

anthers 1-1.5 mm.

not saccate basally;

lower glumes usually 1/5 – 2/3 as long as the spikelets, occasionally equaling the upper florets, (0)1-11-veined;

upper glumes 5-13-veined;

lower florets sterile or staminate;

lower lemmas similar to the upper glumes, 5-9-veined;

lower paleas if present, usually hyaline, 2-veined;

upper florets bisexual, sessile, ovoid to ellipsoid, usually plano-convex, usually glabrous, not disarticulating, mucronate or acuminate;

upper lemmas indurate, transversely rugose and verrucose, 5-veined, margins involute, apices round to mucronate, or aristate;

upper palea rugose, shiny or lustrous;

lodicules 2, cuneate, truncate;

anthers 3.

Caryopses

1.8-2 mm.

ovoid to elliptic, dorsally compressed;

embryos 1/2 - 3/4 as long as the caryopses;

hila punctate to linear, x = 7, 8, 9, or 10.

Lower

sheaths with papillose-based hairs, these more dense distally, margins ciliate;

collars pubescent;

ligules 1-1.5 mm;

blades 7.5-35 cm long, 4-20 mm wide, glabrous or sparsely pilose on both surfaces, margins scabrous.

2n

= 18, 36.

Urochloa mutica

Urochloa

Distribution
from FNA
AL; FL; MD; OR; SC; TX; HI; PR; Virgin Islands
[BONAP county map]
from FNA
AL; AR; AZ; CA; FL; GA; IL; KS; KY; LA; MA; MD; MO; MS; NC; NJ; NM; OK; OR; PA; SC; TN; TX; VA; HI; PR; Virgin Islands
[BONAP county map]
Discussion

An African species, Urochloa mutica is grown as a forage crop throughout the tropics, but it tends to become weedy. It grows on moist, disturbed soils and is established in the southeastern United States.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Urochloa is a genus of approximately 100 tropical and subtropical species. There are 19 species found in the Flora region. Eight species are established introductions, six are native, three are cultivated as grain or forage crops, and two have been found in the region but are not known to be established.

Urochloa differs from Brachiaria in its two or more ranks of panicles and rugose, non-disarticulating, distal florets. The rugose, often mucronate or aristate, distal florets also distinguish it from most species of Panicum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Branches at the lowest node of the inflorescence verticillate; inflorescences simple, open panicles, the primary branches with well-developed secondary and tertiary branches; plants perennial
M. maximus
1. Branches at the lowest node of the inflorescence not verticillate; inflorescences usually with spikelike branches, secondary branches absent or inconspicuous, sometimes simple panicles; plants annual or perennial.
→ 2
2. Spikelets paired at mid-branch length, sometimes solitary distally.
→ 3
3. Axes of the primary panicle branches flat; lower glumes (0)1-3-veined.
→ 4
4. Plants annual; culms 10-35 cm tall; spikelets 1.8-2.2 mm long
U. reptans
4. Plants perennial; culms 20-500 cm tall; spikelets 2.5-5 mm long.
→ 5
5. Upper lemmas awned, the awns 0.5-1.2 mm; lower glumes 3-veined, with 1-3 conspicuous, rigid hairs
U. mosambicensis
5. Upper lemmas unawned; lower glumes 1-3-veined, glabrous
U. mutica
3. Axes of the primary panicle branches triquetrous; lower glumes 3-7-veined.
→ 6
6. Spikelets 4.8-6 mm long; hila linear, about 1/2 as long as the caryopses
U. texana
6. Spikelets 2-4.2 mm long; hila punctate.
→ 7
7. Branch axes densely hairy, the hairs papillose-based
U. arizonica
7. Branch axes sometimes densely hairy, with few or no papillose-based hairs.
→ 8
8. Upper glumes with evident cross venation extending from near the bases to the apices; spikelets obovoid; upper glumes and lower lemmas usually glabrous; lower lemmas 7-veined
U. fusca
8. Upper glumes without evident cross venation, or the cross venation confined to the distal 1/2; spikelets ellipsoid; upper glumes and lower lemmas glabrous or pubescent; lower lemmas 5-veined.
→ 9
9. Cauline nodes glabrous; plants 20-120 cm tall; spikelet apices abruptly acuminate
U. adspersa
9. Cauline nodes pubescent; plants 10-70 cm tall, spikelets apices broadly acute to acute.
→ 10
10. Upper glumes 5-veined; upper lemmas 1.8-2.1 mm long; glumes separated by an internode about 0.3 mm long
U. villosa
10. Upper glumes 7-9-veined; upper lemmas 2.3-3.3 mm long; glumes not separated by a conspicuous internode
U. ramosa
2. Spikelets solitary at mid-branch length.
→ 11
11. Panicle branches triquetrous, 0.2-0.4 mm wide.
→ 12
12. Plants perennial; upper glumes (9) 11-13-veined; upper lemmas 2.4-2.8 mm long; glumes not separated by a conspicuous internode; lower florets staminate
U. ciliatissima
12. Plants annual; upper glumes 5-veined; upper lemmas 1.8-2.1 mm long; glumes separated by an internode about 0.3 mm long; lower florets sterile
U. villosa
11. Panicle branches flat or crescent-shaped in cross section, 0.5-2.5 mm wide.
→ 13
13. Upper lemmas awned, the awns 0.3-1.2 mm long, apices rounded.
→ 14
14. Plants perennial; lower glumes with 1-3 conspicuous, rigid hairs, the lower glumes 1/2 - 3/4 as long as the spikelet; lower florets staminate
U. mosambicensis
14. Plants annual; lower glumes without conspicuous, rigid hairs; the lower glumes 1/4 - 1/3(1/2) as long as the spikelet; lower florets sterile
U. panicoides
13. Upper lemmas unawned; apices variable, with or without a mucronate tip.
→ 15
15. Spikelets in a single row along the branches; spikelets 4-6 mm long; lower florets staminate; panicle branch axes crescent-shaped in cross section
U. brizantha
15. Spikelets in 2 rows along the branches; spikelets 2.5-6 mm long; lower florets sterile or staminate; panicle branches flat.
→ 16
16. Lower glumes 5-7-veined; glumes scarcely separated, the internode between them shorter than 0.3 mm.
→ 17
17. Upper glumes and lower lemmas pubescent, the hairs often long in the distal 1/3
U. piligera
17. Upper glumes and lower lemmas glabrous.
→ 18
18. Plants perennial, stoloniferous; lower florets staminate
U. arrecta
18. Plants annual; lower florets sterile
U. platyphylla
16. Lower glumes (7)9-11-veined; glumes separated by a conspicuous, 0.3-0.5 mm internode.
→ 19
19. Lower paleas absent; spikelets usually pubescent, sometimes glabrous
U. piligera
19. Lower paleas present; spikelets glabrous.
→ 20
20. Spikelets 3.3-3.7 mm long; base of blades rounded to subcordate, not clasping the stem
U. subquadripara
20. Spikelets 4-6 mm long; base of blades subcordate to cordate, clasping the stem
U. plantaginea
Source FNA vol. 25, p. 494. FNA vol. 25, p. 492. Author: J.K. Wipff; Rahmona A. Thompson;.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Urochloa Poaceae > subfam. Panicoideae > tribe Paniceae
Sibling taxa
U. adspersa, U. arizonica, U. arrecta, U. brizantha, U. ciliatissima, U. fusca, U. mosambicensis, U. panicoides, U. piligera, U. plantaginea, U. platyphylla, U. ramosa, U. reptans, U. subquadripara, U. texana, U. villosa
Subordinate taxa
M. maximus, U. adspersa, U. arizonica, U. arrecta, U. brizantha, U. ciliatissima, U. fusca, U. mosambicensis, U. mutica, U. panicoides, U. piligera, U. plantaginea, U. platyphylla, U. ramosa, U. reptans, U. subquadripara, U. texana, U. villosa
Synonyms Panicum purpurascens, Brachiaria mutica
Name authority (Forssk.) T.Q. Nguyen P. Beauv.
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