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nodding false oat, nodding oatgrass, nodding trisetum

false oat, oatgrass, trisetum

Habit Plants perennial, with both fertile and sterile shoots; cespitose, not rhizomatous. Plants annual or perennial; sometimes rhizomatous, sometimes cespitose.
Culms

(30)50-110 cm, clumped, erect, glabrous or pubescent.

5-150 cm, glabrous or pubescent, basal branching extravaginal.

Sheaths

open the entire length or fused at the base;

auricles absent;

ligules membranous, often erose to lacerate, sometimes ciliolate;

blades rolled in the bud.

Leaves

2-3 per culm;

sheaths scabridulous or pilose;

ligules 1.5-3 mm, truncate, erose to lacerate;

blades (8.5)15-20+ cm long, (3)7-12 mm wide, flat, ascending, lax at maturity, often scabridulous.

Panicles

10-30 cm long, (1)2-9 cm wide, open, nodding, green or tan, occasionally purple-tinged;

branches 2-12+ cm, most, except sometimes the uppermost, spikelet-bearing only towards the apices, with the basal (1/5)1/3-1/2 bare, filiform, flexuous, at least the lowest 1-3 whorls spreading or drooping.

Inflorescences

terminal panicles, open and diffuse to dense and spikelike;

branches antrorsely scabrous.

Spikelets

6-12 mm, subsessile to pedicellate, pedicels to 2 cm, usually with 2-3 functional florets below 1-2 reduced florets;

rachilla internodes and hairs 1-2.5 mm;

disarticulation above the glumes, beneath the florets.

2.5-12 mm, usually subsessile to pedicellate, rarely sessile, laterally compressed, with 2-5 florets, reduced florets sometimes present distally;

rachillas hairy, internodes evident, prolonged beyond the base of the distal bisexual florets;

disarticulation usually initially above the glumes and beneath the florets, subsequently below the glumes, in some species initially below the glumes.

Glumes

unequal;

lower glumes 0.75-2(3) mm, subulate;

upper glumes 3.5-5 mm long, shorter than the lowest florets, 2-3 times as wide as the lower glumes, widest at or above the middle, ovate or obovate, rounded to the acuminate apices;

callus hairs to 1 mm;

lemmas 5-6 mm, broadly lanceolate, glabrous, bifid, teeth to 1.3 mm, awned, awns (7)9-14 mm, arising from above midlength to just below the teeth, exceeding the lemma apices, arcuate to flexuous;

paleas shorter than the lemmas;

anthers about 1 mm.

subequal or unequal, keels scabrous, apices usually acute, unawned, often apiculate;

lower glumes 1(3)-veined;

upper glumes 3(5)-veined, lateral veins less than 1/2 the glume length;

calluses hairy;

lemmas 3-7-veined, margins hyaline, unawned or awned from above the middle with a single awn, apices usually bifid, sometimes entire;

paleas from subequal to longer than the lemmas, membranous, 2-veined, veins usually extending as bristlelike tips;

lodicules 2, shallowly and usually slenderly lobed to fimbriate;

anthers 3;

ovaries glabrous or pubescent;

styles 2.

Caryopses

2.5-3.2 mm, densely to sparsely pubescent.

shorter than the lemmas, concealed at maturity, elongate-fusiform, compressed, brown;

embryos elliptic, to 1/3 the length of the caryopses;

endosperm milky, x = 7.

2n

= 42.

Trisetum cernuum

Trisetum

Distribution
from USDA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; CT; ID; KS; LA; MA; ME; MI; MN; MO; MS; MT; NC; NH; NJ; NM; NV; NY; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WY; HI; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Greenland
[BONAP county map]
Discussion

Trisetum cernuum grows in moist woods, on stream banks, lake and pond shores, and floodplains of the western Flora region. The hairiness of the leaf sheaths varies, often within a plant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trisetum, a genus of approximately 75 species, occurs primarily in temperate, subarctic, and alpine regions. Eight species are native to the Flora region; two have been introduced, one of which is not known to have persisted. Trisetum usually differs from Sphenopholis in having longer awns that are inserted lower on the lemmas, and spikelets that disarticulate above the glumes. It differs from Deschampsia primarily in its more acute, bifid lemmas. In addition, all species of Trisetum have awns that are inserted at or above the midpoint of the lemmas; in Deschampsia, the awns are usually inserted at or below midlength, often near the base.

Trisetum spicatum is important as forage on native rangelands. Like other species of the genus, it is a significant component of natural food pyramids, especially in arctic and alpine regions and mountain parks. Trisetum flavescens was introduced from Europe as a pasture grass; T. interruptum is often weedy.

Spikelet measurements do not include the awns.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants annual; without sterile shoots.
→ 2
2. Lower glumes 3-veined; spikelets 3-6 mm long; panicles 0.3-1.5 cm wide; plants native
T. interruptum
2. Lower glumes 1-veined; spikelets 2.5-3.5 mm long; panicles 0.5-3 cm wide; plants not native, not established
T. aureum
1. Plants perennial; usually producing both fertile and sterile shoots.
→ 3
3. Lemmas unawned or with inconspicuous straight awns up to 2 mm long that rarely exceed the lemma apices.
→ 4
4. Panicles usually 2-4 cm wide, lax and nodding; callus and rachilla hairs 1.3-2 mm long; plants of eastern North America
T. melicoides
4. Panicles usually 1-1.5 cm wide, erect; callus hairs shorter than 0.5 mm; rachilla hairs up to 1 mm long; plants of western North America
T. wolfii
3. Lemmas with evident awns 3-14 mm long, these straight, curved, flexuous, or geniculate, exceeding the lemma apices.
→ 5
5. Plants rhizomatous; culms usually solitary.
→ 6
6. Lemma teeth usually 3-6 mm long; ligules 0.5-1(2) mm long
T. flavescens
6. Lemma teeth usually shorter than 1 mm; ligules 1-5 mm long.
→ 7
7. Culms 15-65 cm tall; panicles 2-12(16) cm long; plants of Alaska and the Yukon Territory
T. sibiricum
7. Culms 80-110 cm tall; panicles 13-20 cm long; known only from Montana
T. orthochaetum
5. Plants not rhizomatous; culms clumped.
→ 8
8. Glumes usually subequal; both glumes lanceolate; upper glumes less than twice as wide as the lower glumes
T. spicatum
8. Glumes usually unequal, sometimes subequal; lower glumes subulate to linear-lanceolate or lance-elliptic; upper glumes broadly lanceolate to ovate or obovate, at least twice as wide as the lower glumes.
→ 9
9. Upper glumes as long as or longer than the lowest florets; awns 3-9 mm long; rachilla hairs up to 1.5 mm long; ligules 0.5-2 mm long; panicles yellowish brown
T. flavescens
9. Upper glumes shorter than the lowest florets; awns 7-14 mm long; rachilla hairs 0.7-2.5 mm long; ligules 1.5-6 mm long; panicles green or tan.
→ 10
10. Most panicle branches, except sometimes the lowermost, spikelet-bearing for their full length; panicles erect or nodding at the apices; branches ascending to somewhat divergent; upper glumes widest at or below the middle, tapering to the apices; lower glumes 3-5 mm long
T. canescens
10. Most panicle branches, except sometimes the uppermost, spikelet-bearing only towards the apices; panicles nodding; branches of at least the lowest 1-3 whorls spreading or drooping; upper glumes widest at or above the middle, rounded to the apices; lower glumes 0.75-3 mm long
T. cernuum
Source FNA vol. 24, p. 748. FNA vol. 24, p. 744. Author: John H. Rumely;.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Trisetum Poaceae > subfam. Pooideae > tribe Poeae
Sibling taxa
T. aureum, T. canescens, T. flavescens, T. interruptum, T. melicoides, T. orthochaetum, T. sibiricum, T. spicatum, T. wolfii
Subordinate taxa
T. aureum, T. canescens, T. cernuum, T. flavescens, T. interruptum, T. melicoides, T. orthochaetum, T. sibiricum, T. spicatum, T. wolfii
Synonyms T. nutkanense
Name authority Trin. Pers.
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