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little sand verbena, sandpuffs, small-flower sand-verbena, small-flower sandpuffs

sand -puffs

Habit Herbs, annual, viscid pubescent to nearly glabrous, from slender or stout and ± woody taproots.
Stems

reddish, glandular-pubescent, ± viscid.

decumbent to semierect, unarmed, without glutinous bands on internodes.

Leaves

petiole 1–4 cm;

blade lance-ovate to elliptic, 1–6 × 0.5–2.5 cm, margins entire or sinuate, ciliate;

surfaces short glandular-pubescent, ± viscid, abaxial surface green or glaucous, adaxial surface green.

petiolate, unequal in size in each pair;

blade ± thick and succulent, base usually asymmetric.

Inflorescences

5–15-flowered;

bracts lanceolate to ovate, 3–9 × 1–3 mm, thin and green or ± papery, base ± attenuate, margins glandular-puberulent, often ciliate, or glabrate to lightly glandular-puberulent.

axillary, pedunculate, capitate clusters;

receptacle flat to somewhat rounded-conic, with short, pedicel-like projections;

flowers maturing from one side of inflorescence to other;

bracts persistent, not accrescent, 5–10, distinct, thin and translucent, forming an involucre, linear-lanceolate to ovate, broad.

Flowers

bisexual, chasmogamous;

perianth radially symmetric, funnelform or salverform, constricted beyond ovary, abruptly expanded to 4–5-lobed limb;

stamens (3–)4–5, included;

styles included;

stigmas linear.

Perianth

tube greenish to pink, 6–18 mm, limb greenish to pink, 3–5 mm diam., lobes inconspicuous.

Fruits

oval to round in profile, 10–20 × 10–20 mm, walls ± spongy on exterior, indurate on interior, ± smooth to rugose;

lateral ribs, if extending into wings, only slightly raised;

wings (2–)3(–4).

fusiform, indurate throughout, or spongy on exterior, minutely puberulent or glabrous;

wings 2–4, translucent, prominently veined, scarious, extending beyond apex and/or base of body;

sulci smooth or coarsely rugose.

Tripterocalyx micranthus

Tripterocalyx

Phenology Flowering spring–fall.
Habitat Sandy soils, desert scrub, desert grasslands
Elevation 800-2400 m (2600-7900 ft)
Distribution
from FNA
AZ; CA; CO; KS; MT; ND; NM; NV; SD; UT; WY; AB; SK
[WildflowerSearch map]
[BONAP county map]
from USDA
North America (including n Mexico)
[BONAP county map]
Discussion

The names Abronia cycloptera A. Gray and Tripterocalyx cyclopterus (A. Gray) Standley are illegitimate, based on an attempt by Gray to rename A. micrantha Torrey. Gray believed A. micrantha to be a “precociously fertilized state” of what is here recognized at T. carneus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits usually with 2 wings, body occasionally ribbed laterally, extending prominently into wings
T. crux-maltae
1. Fruits usually with more than 2 wings, body often ribbed laterally, lateral ribs, if extending into wings, only slightly raised
→ 2
2. Perianth tube 6-18 mm; perianth limb 3-5 mm diam., lobes inconspicous
T. micranthus
2. Perianth tube 12-30 mm; perianth limb 8-13 mm diam., lobes over 1.5 mm, showy
→ 3
3. Perianth limb pale pink to magenta; fruits usually over 20 mm
T. carneus
3. Perianth limb white adaxially, white to pink abaxially; fruits usually 20 mm or less
T. wootonii
Source FNA vol. 4, p. 70. FNA vol. 4, p. 70. Author: Leo A. Galloway.
Parent taxa Nyctaginaceae > Tripterocalyx Nyctaginaceae
Sibling taxa
T. carneus, T. crux-maltae, T. wootonii
Subordinate taxa
T. carneus, T. crux-maltae, T. micranthus, T. wootonii
Synonyms Abronia micrantha Abronia unranked T.
Name authority (Torrey) Hooker: Hooker’s J. Bot. Kew Gard. Misc. 5: 261. (1853) (Torrey) Hooker: Hooker’s J. Bot. Kew Gard. Misc. 5: 261. (1853)
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