Trillium maculatum
Trillium luteum
spotted trillium, spotted wakerobin
wax trillium, yellow toadshade, yellow trillium, yellow wakerobin
horizontal, brownish, short, thick, praemorse, not brittle.
horizontal, brownish, short, thick, praemorse, not brittle.
1–3, round in cross section, 1.4–4 dm, slender to stout, glabrous.
1–4, round in cross section, 1.4–4 dm, stout, glabrous.
held well above ground, tips only touching ground in early anthesis, sessile;
blade dark green, mottling darker, mottled to varying degrees, mottling becoming obscure with age, broadly ovate-elliptic to elliptic, 7–15 × 4.6–6.7 cm, not glossy, apex rounded or barely acuminate.
held well above ground, sessile;
blade mottled, mottling becoming more obscure as growth matures, shape variable, ovate-elliptic to ± orbicular-acuminate, 6.5–17 × 6.5–9.8 cm, not glossy, apex acuminate to long-acuminate.
erect, odor faintly spicy and bananalike;
sepals displayed above bracts, spreading almost to horizontal, often suffused or streaked with purple-maroon, lanceolate-linear, 22–50 × 5–7 mm, margins entire, apex rounded-acute, recurving slightly;
petals long-lasting, spreading-erect, ± connivent, ± concealing stamens and ovary or sometimes slightly spreading early, only partially obscuring stamens, becoming more erect and even touching in the manner of Trillium cuneatum when older, clear dark red-maroon, or reddish purple, without muddy or brown tones of many purple sessile species, or, rarely, purple basally and yellow toward apex, or clear sulfur yellow, not spirally twisted, narrowly oblanceolate-spatulate to linear-spatulate, widest above middle, narrowing to apex, 4–7 × 0.7–1.7 cm, thick-textured, margins entire, slightly involute in proximal 1/2, apex acute to round-acute;
stamens erect, 12–20 mm;
filaments dark purple, 2–3 mm, widest at base, much shorter than anther sacs;
anthers erect, straight, brownish purple, 10–16 mm, dehiscence introrse on broad connective;
connectives brown-purple, straight, essentially not extended beyond anther sacs;
ovary dark purple, ovoid, weakly 3-angled to smooth (rarely very obscurely 6-angled), angles often obscured in large, turgid ovary, 8–11 mm;
stigmas erect, divergent-recurved, distinct, purplish, subulate, 2–4 mm, fleshy.
erect, odor strongly lemony;
sepals displayed above bracts, widely spreading, green, oblong-lanceolate, lanceolate, or oblong-elliptic, 22–47 × 8–9 mm, margins entire, apex distinctly rounded-acute;
petals long-lasting, erect, ± connivent, ± concealing stamens and ovary, greenish yellow to lemon yellow, not spirally twisted, not clawed, elliptic-lanceolate, lanceolate, or oblanceolate, 3.4–6.6 × 1–2.1 cm, thick-textured, widest near base, margins entire, tapering to long-acuminate apex;
stamens erect, 11–18 mm;
filaments greenish white, 1.5–2 mm, basally dilated;
anthers erect, straight, yellow, 9–16 mm, dehiscence introrse (or appearing lateral);
connectives green, straight, broad, extending only slightly (to 0.5 mm) or not at all beyond anther sacs;
ovary pale green, ovoid to globose, 6-angled, 5–8 mm;
stigmas erect, barely spreading, distinct, greenish white, sessile, linear to subulate, 3–4 mm, fleshy.
dark purplish green, odor not reported, ovoid, obscurely 3- to 6-angled, bearing persistent stigma, 1 × 2 cm, pulpy.
green to greenish white, occasionally with dark streaks, odor not reported, ovoid, 6-angled, 3.5 × 2 cm, pulpy.
= 10.
Trillium maculatum
Trillium luteum
In their account of the Carolinas flora, A. E. Radford et al. (1968) did not treat Trillium maculatum or give any explanation of why they omitted this species, which is fairly widely distributed in southeastern South Carolina and morphologically is one of the more distinct sessile trilliums.
Many color forms occur and have been named. In forma luteum J. D. Freeman, for example, all floral organs lack purple pigment, and the petals are clear, soft yellow, whereas forma simulans J. D. Freeman has yellow petals with purple bases, and purple stamens and carpels. These are illustrated in color in F. W. Case and R. B. Case (1997).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Botanists have been confused by Trillium luteum for a long time. Some, such as A. E. Radford et al. (1968), appear to regard it as a form of T. cuneatum, while others confuse it with T. viride, a more western species. Early botanists confused T. luteum with the occasional individual or very local larger population of pallid color forms of other species. Trillium cuneatum rather frequently produces green, yellowish green, or pale lemon yellow forms (but with a cuneate larger and wider petal) that mimic T. luteum. These forms, when growing with T. luteum, hybridize, leading to so many intergrades that many plants cannot be placed in either species with any confidence. For these reasons, almost no work older than J. D. Freeman’s (1975) can be used reliably to plot distribution of T. luteum.
In some parts of their ranges, Trillium luteum and T. cuneatum do not grow together. Trillium luteum is abundant in eastern Tennessee, extending a short distance into North Carolina and Georgia. Trillium cuneatum is rare or absent from this region except along the Little Tennessee River (J. D. Freeman 1975).
Trillium luteum has escaped and become established along the Grand and Saint Joseph rivers and elsewhere in Michigan, near Kingston and Grimsby, Ontario (J. K. Morton pers. comm.), and probably elsewhere.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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