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giant purple wakerobin

spotted trillium, spotted wakerobin

Rhizomes

horizontal, brownish, thick, praemorse, not brittle.

horizontal, brownish, short, thick, praemorse, not brittle.

Scapes

often 2 from single terminal bud, round in cross section, 2.5–5.5 dm, stout, glabrous.

1–3, round in cross section, 1.4–4 dm, slender to stout, glabrous.

Bracts

held well above ground, sessile;

blade bright green (in early anthesis rather succulent in appearance), usually well-marked with lighter and darker green spots, occasionally obscurely or scarcely mottled, mottling becoming obscure with age, ovate to broadly ovate, 11–18 × 12–17 cm, somewhat glossy, apex acuminate.

held well above ground, tips only touching ground in early anthesis, sessile;

blade dark green, mottling darker, mottled to varying degrees, mottling becoming obscure with age, broadly ovate-elliptic to elliptic, 7–15 × 4.6–6.7 cm, not glossy, apex rounded or barely acuminate.

Flower

odor spicy in fresh flowers, becoming fetid in older flowers;

sepals displayed above bracts, spreading, divergent, green, purple-streaked, lanceolate, 40–75 × 10–14 mm, margins entire, apex acute to round-acute;

petals long-lasting, conspicuous, spreading-erect to erect, tips incurving slightly, ± connivent, partially to completely concealing stamens and ovary, glossy dark maroon-red or purple, not spirally twisted, flat, oblanceolate, 5.5–11 × 2–3.5 cm, widest at or below middle, glossy, thick-textured, cuneate basally, margins ± flat, entire, apex round-acute;

stamens erect, straight, 15–26 mm, slightly concealing ovary;

filaments dark purple, 2–4 mm;

anthers dark maroon, 13–24 mm, dehiscence introrse;

connectives straight, barely extended beyond anther sacs;

ovary inconspicuous, purple, ovoid, round to 6-angled, 8–15 mm;

stigmas erect, distinct, subulate, 6–8 mm, fleshy, thickened basally.

erect, odor faintly spicy and bananalike;

sepals displayed above bracts, spreading almost to horizontal, often suffused or streaked with purple-maroon, lanceolate-linear, 22–50 × 5–7 mm, margins entire, apex rounded-acute, recurving slightly;

petals long-lasting, spreading-erect, ± connivent, ± concealing stamens and ovary or sometimes slightly spreading early, only partially obscuring stamens, becoming more erect and even touching in the manner of Trillium cuneatum when older, clear dark red-maroon, or reddish purple, without muddy or brown tones of many purple sessile species, or, rarely, purple basally and yellow toward apex, or clear sulfur yellow, not spirally twisted, narrowly oblanceolate-spatulate to linear-spatulate, widest above middle, narrowing to apex, 4–7 × 0.7–1.7 cm, thick-textured, margins entire, slightly involute in proximal 1/2, apex acute to round-acute;

stamens erect, 12–20 mm;

filaments dark purple, 2–3 mm, widest at base, much shorter than anther sacs;

anthers erect, straight, brownish purple, 10–16 mm, dehiscence introrse on broad connective;

connectives brown-purple, straight, essentially not extended beyond anther sacs;

ovary dark purple, ovoid, weakly 3-angled to smooth (rarely very obscurely 6-angled), angles often obscured in large, turgid ovary, 8–11 mm;

stigmas erect, divergent-recurved, distinct, purplish, subulate, 2–4 mm, fleshy.

Fruits

dark reddish purple, ovoid to ellipsoid or weakly angled, 20–50 mm, fleshy.

dark purplish green, odor not reported, ovoid, obscurely 3- to 6-angled, bearing persistent stigma, 1 × 2 cm, pulpy.

Trillium kurabayashii

Trillium maculatum

Phenology Flowering spring (late Mar–early May). Flowering mid winter–early spring (early Feb–early Apr).
Habitat Rich, moist conifer-hardwood forest, slopes, especially lower slopes, predominantly deciduous flat woods along streams, edges of Sequoia groves, and alder, vine maple, and fern thickets along streams, especially older, higher flood terraces, not the lowest and wettest, at higher elevations, both in forests and in open grassy meadows with scattered oak trees Rich mesic forests, particularly banks and bluffs of rivers, floodplains, often where quite brushy, rich soils, calcareous soils, alluvium
Elevation 20–500+ m (100–1600+ ft) 0–500 m (0–1600 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
from FNA
AL; FL; GA; SC
[WildflowerSearch map]
[BONAP county map]
Discussion

In their account of the Carolinas flora, A. E. Radford et al. (1968) did not treat Trillium maculatum or give any explanation of why they omitted this species, which is fairly widely distributed in southeastern South Carolina and morphologically is one of the more distinct sessile trilliums.

Many color forms occur and have been named. In forma luteum J. D. Freeman, for example, all floral organs lack purple pigment, and the petals are clear, soft yellow, whereas forma simulans J. D. Freeman has yellow petals with purple bases, and purple stamens and carpels. These are illustrated in color in F. W. Case and R. B. Case (1997).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 26, p. 110. FNA vol. 26, p. 112.
Parent taxa Liliaceae > Trillium > subg. Phyllantherum Liliaceae > Trillium > subg. Phyllantherum
Sibling taxa
T. albidum, T. angustipetalum, T. catesbaei, T. cernuum, T. chloropetalum, T. cuneatum, T. decipiens, T. decumbens, T. discolor, T. erectum, T. flexipes, T. foetidissimum, T. gracile, T. grandiflorum, T. lancifolium, T. ludovicianum, T. luteum, T. maculatum, T. nivale, T. ovatum, T. parviflorum, T. persistens, T. petiolatum, T. pusillum, T. recurvatum, T. reliquum, T. rivale, T. rugelii, T. sessile, T. simile, T. stamineum, T. sulcatum, T. underwoodii, T. undulatum, T. vaseyi, T. viride, T. viridescens
T. albidum, T. angustipetalum, T. catesbaei, T. cernuum, T. chloropetalum, T. cuneatum, T. decipiens, T. decumbens, T. discolor, T. erectum, T. flexipes, T. foetidissimum, T. gracile, T. grandiflorum, T. kurabayashii, T. lancifolium, T. ludovicianum, T. luteum, T. nivale, T. ovatum, T. parviflorum, T. persistens, T. petiolatum, T. pusillum, T. recurvatum, T. reliquum, T. rivale, T. rugelii, T. sessile, T. simile, T. stamineum, T. sulcatum, T. underwoodii, T. undulatum, T. vaseyi, T. viride, T. viridescens
Name authority J. D. Freeman: Brittonia 27: 56, fig. 12. (1975) Rafinesque: Med. Fl. 2: 103. (1830)
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