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giant purple wakerobin, giant trillium, giant wakerobin, sessile trillium, small-flower trillium

Louisiana trillium, Louisiana wakerobin

Rhizomes

± erect, brownish, somewhat compressed-thickened, superficially bulblike, praemorse, not brittle.

± horizontal, brownish, short, thick, praemorse, not brittle.

Scapes

1–3, green, round in cross section, 2–6.5 dm, robust.

1–3, round in cross section, 1.4–2.6 dm, ± slender, glabrous.

Bracts

held well above ground, sessile (narrowing of bract blade may give bract subsessile appearance);

blade densely to weakly mottled in dark brownish green, mottling becoming more obscure to absent as bract matures, broadly ovate, 7–17.6 × 7.4–17.7 cm, not glossy, apex obtuse-rounded.

held well above ground, sessile;

blade strongly mottled in dark and bronzy green, often with central light strip, mottling becoming somewhat obscure with age, lanceolate-ovate, 5.3–9.5 × 2.3–5 cm, not glossy, margins of distal 1/3 convex-curved to apex, apex rounded-acute.

Flower

erect, odor roselike, spicy;

sepals spreading-ascending above bracts, green, lanceolate, 35–65 × 7–12 mm, margins entire, flat, apex obtusely rounded;

petals long-lasting, erect, connivent, ± concealing stamens and ovary, yellow, bronze, maroon, brown, deep purple, reddish brown, pink, dark purplish red, purplish bronze, rarely greenish white, not spirally twisted, veins not engraved, oblanceolate to obovate, 6.5–10 × 1.5–2.5 cm, thick-textured, base cuneate, margins entire, apex variably acute to almost truncate, erose;

stamens erect, purplish, 17–26 mm;

filaments purple, ca. 4 mm, widest at base, much shorter than anther sacs;

anthers erect, straight, ± purple-brown, 13–22 mm, dehiscence introrse;

connectives purple, straight, extended ca. 1–1.5 mm beyond anther sacs;

ovary purple, ovoid, 6-angled, 6–12 mm;

stigmas small, divergent or erect, distinct, purple, subulate, 4–8 mm, not fleshy.

erect, odor of carrion;

sepals displayed above bracts, spreading, green, lanceolate-oblanceolate, 19–35 × 2.7–4 mm, margins entire, apex rounded or acute to sometimes weakly reflexed;

petals long-lasting, faintly introrsely curved-erect-spreading, weakly connivent, ± concealing stamens and ovary, dark maroon-brown, purplish, or dull greenish, or 2-colored, basal portion purple, distal portion grayish green, not spirally twisted, oblanceolate-linear, 3.5–5.5 × 0.4–0.8 cm, thick-textured, thickened and weakly clawed basally, margins entire, apex acute;

stamens erect, straight, 10–18 mm;

filaments olive-orange, 2–3 mm, widened basally;

anthers erect, straight, olive to orange, 7–20 mm, slender, dehiscence latrorse;

connectives olive to orange, straight, scarcely extended beyond anther sac;

ovary purple, ovoid, 6-angled, 8–9 mm;

stigmas erect, with spreading or coiled tips, distinct, pale purple, subulate, 3–6 mm, ± fleshy.

Fruits

red-purple, fragrance not reported, ovoid, obscurely 6-angled, 2.5–3 cm, pulpy, juicy.

dark purplish green, little or no odor, ovoid, 6-angled, pulpy.

2n

= 10.

Trillium chloropetalum

Trillium ludovicianum

Phenology Flowering late winter–early spring (early Mar–Apr).
Habitat Low flatwoods, floodplains along streams, steep ravine slopes leading to floodplains, mixed pine-beech woods
Elevation 50–500 m (200–1600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
LA; MS
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

J. D. Freeman (1975) considered that Trillium chloropetalum differs from T. albidum in having introrse (not latrorse) anther sacs, and that the purple pigments present on anther and ovary tissue here are absent in T. albidum. In some places, hybridization between the two certainly has occurred, and a complete range of intergrades exists.

This species merits further study. The following varieties are only weakly differentiated and perhaps ought to be dropped.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The range of Trillium ludovicianum is near to that of T. cuneatum in Mississippi, and the two appear to intergrade.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petal tissue always with yellow pigments, often present with other pigments; petals yellow, greenish yellow, greenish purple, or bronze-green or brown.
var. chloropetalum
1. Petal tissue lacking yellow pigments; petals purple, reddish purple, garnet red, pink, or greenish white.
var. giganteum
Source FNA vol. 26. FNA vol. 26, p. 111.
Parent taxa Liliaceae > Trillium > subg. Phyllantherum Liliaceae > Trillium > subg. Phyllantherum
Sibling taxa
T. albidum, T. angustipetalum, T. catesbaei, T. cernuum, T. cuneatum, T. decipiens, T. decumbens, T. discolor, T. erectum, T. flexipes, T. foetidissimum, T. gracile, T. grandiflorum, T. kurabayashii, T. lancifolium, T. ludovicianum, T. luteum, T. maculatum, T. nivale, T. ovatum, T. parviflorum, T. persistens, T. petiolatum, T. pusillum, T. recurvatum, T. reliquum, T. rivale, T. rugelii, T. sessile, T. simile, T. stamineum, T. sulcatum, T. underwoodii, T. undulatum, T. vaseyi, T. viride, T. viridescens
T. albidum, T. angustipetalum, T. catesbaei, T. cernuum, T. chloropetalum, T. cuneatum, T. decipiens, T. decumbens, T. discolor, T. erectum, T. flexipes, T. foetidissimum, T. gracile, T. grandiflorum, T. kurabayashii, T. lancifolium, T. luteum, T. maculatum, T. nivale, T. ovatum, T. parviflorum, T. persistens, T. petiolatum, T. pusillum, T. recurvatum, T. reliquum, T. rivale, T. rugelii, T. sessile, T. simile, T. stamineum, T. sulcatum, T. underwoodii, T. undulatum, T. vaseyi, T. viride, T. viridescens
Subordinate taxa
T. chloropetalum var. chloropetalum, T. chloropetalum var. giganteum
Synonyms T. sessile var. chloropetalum, T. giganteum var. chloropetalum
Name authority (Torrey) Howell: Fl. N.W. Amer., 661. (1902) Harbison: Biltmore Bot. Stud. 1: 23. (1901)
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