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giant purple wakerobin, giant trillium, giant wakerobin, sessile trillium, small-flower trillium

giant purple wakerobin, narrow petal wakerobin, narrow-petal trillium

Habit Plants tall, very robust.
Rhizomes

± erect, brownish, somewhat compressed-thickened, superficially bulblike, praemorse, not brittle.

erect, brownish, thick, somewhat compressed-thickened, praemorse, not brittle.

Scapes

1–3, green, round in cross section, 2–6.5 dm, robust.

1–2, round in cross section, 2.5–6 dm.

Bracts

held well above ground, sessile (narrowing of bract blade may give bract subsessile appearance);

blade densely to weakly mottled in dark brownish green, mottling becoming more obscure to absent as bract matures, broadly ovate, 7–17.6 × 7.4–17.7 cm, not glossy, apex obtuse-rounded.

held well above ground, spreading horizontally, subsessile;

blade very sparsely mottled with dark greenish brown or rarely all green, mottling becoming obscure with age, broadly ovate, 10–22 × 8.7–15 cm, not glossy, often narrowed to falsely petiolate, very short, and narrowly cuneate base 10–20 mm, apex obtuse.

Flower

erect, odor roselike, spicy;

sepals spreading-ascending above bracts, green, lanceolate, 35–65 × 7–12 mm, margins entire, flat, apex obtusely rounded;

petals long-lasting, erect, connivent, ± concealing stamens and ovary, yellow, bronze, maroon, brown, deep purple, reddish brown, pink, dark purplish red, purplish bronze, rarely greenish white, not spirally twisted, veins not engraved, oblanceolate to obovate, 6.5–10 × 1.5–2.5 cm, thick-textured, base cuneate, margins entire, apex variably acute to almost truncate, erose;

stamens erect, purplish, 17–26 mm;

filaments purple, ca. 4 mm, widest at base, much shorter than anther sacs;

anthers erect, straight, ± purple-brown, 13–22 mm, dehiscence introrse;

connectives purple, straight, extended ca. 1–1.5 mm beyond anther sacs;

ovary purple, ovoid, 6-angled, 6–12 mm;

stigmas small, divergent or erect, distinct, purple, subulate, 4–8 mm, not fleshy.

erect, odor spicy-musty, musty, or fetid;

sepals conspicuous, spreading, often resting on bracts, maroon to green, linear to oblong-lanceolate, 35–47 × 8–10 mm, margins flat, entire, apex acute;

petals long-lasting, erect, ± connivent, ± concealing stamens and ovary and partially obscuring stamens, dark purple to red-purple, not spirally twisted, veins obscure, linear, 5–10 × 0.7–1.4 cm, 8–10 times longer than wide, glossy, thick-textured, base linear, margins entire, at first flat but inrolling with age, apex variously acute-obtuse;

stamens erect, 12–22 mm;

filaments dark maroon, 2–4 mm, slender, widest at base;

anthers erect, straight, purple, 12–18 mm, dehiscence introrse;

connectives purple, slightly extended 1–1.5 mm beyond anther sacs;

ovary dark, ovoid-ellipsoid, 6-angled toward apex, 7.5–12 mm;

stigmas erect, divergent-recurved, distinct, purple, sessile, awl-shaped, thickly subulate, 5 mm, thick, fleshy.

Fruits

red-purple, fragrance not reported, ovoid, obscurely 6-angled, 2.5–3 cm, pulpy, juicy.

dark purple, fragrance unknown, subglobose, 6-angled, almost winged, fleshy.

2n

= 10.

= 10.

Trillium chloropetalum

Trillium angustipetalum

Phenology Flowering spring–early summer (Mar–Jun).
Habitat Big-tree (Sequoiadendron) groves and other mixed coniferous-deciduous flatwoods, slightly damper depressions under maples and deciduous shrubs, coastal mountains, oak (Quercus) groves in ravines and otherwise quite arid, almost treeless chaparral, wooded canyon slopes, dense woods near streams
Elevation 30–200 m (100–700 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

J. D. Freeman (1975) considered that Trillium chloropetalum differs from T. albidum in having introrse (not latrorse) anther sacs, and that the purple pigments present on anther and ovary tissue here are absent in T. albidum. In some places, hybridization between the two certainly has occurred, and a complete range of intergrades exists.

This species merits further study. The following varieties are only weakly differentiated and perhaps ought to be dropped.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trillium angustipetalum occurs in the Sierra Nevada from Fresno County north to Placer County (J. D. Freeman 1975). It is disjunct in the coastal mountains and hills of Santa Barbara and San Luis Obispo counties.

B. D. Ness (1993) listed Trillium kurabayashii as a synonym of T. angustipetalum. In bract orientation, color, and texture, and in petal shape, the two are quite different and certainly not the same species. Cytologist Masataka Kurabayashi found chromosomal differences between the two species (reported by J. D. Freeman 1975).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petal tissue always with yellow pigments, often present with other pigments; petals yellow, greenish yellow, greenish purple, or bronze-green or brown.
var. chloropetalum
1. Petal tissue lacking yellow pigments; petals purple, reddish purple, garnet red, pink, or greenish white.
var. giganteum
Source FNA vol. 26. FNA vol. 26.
Parent taxa Liliaceae > Trillium > subg. Phyllantherum Liliaceae > Trillium > subg. Phyllantherum
Sibling taxa
T. albidum, T. angustipetalum, T. catesbaei, T. cernuum, T. cuneatum, T. decipiens, T. decumbens, T. discolor, T. erectum, T. flexipes, T. foetidissimum, T. gracile, T. grandiflorum, T. kurabayashii, T. lancifolium, T. ludovicianum, T. luteum, T. maculatum, T. nivale, T. ovatum, T. parviflorum, T. persistens, T. petiolatum, T. pusillum, T. recurvatum, T. reliquum, T. rivale, T. rugelii, T. sessile, T. simile, T. stamineum, T. sulcatum, T. underwoodii, T. undulatum, T. vaseyi, T. viride, T. viridescens
T. albidum, T. catesbaei, T. cernuum, T. chloropetalum, T. cuneatum, T. decipiens, T. decumbens, T. discolor, T. erectum, T. flexipes, T. foetidissimum, T. gracile, T. grandiflorum, T. kurabayashii, T. lancifolium, T. ludovicianum, T. luteum, T. maculatum, T. nivale, T. ovatum, T. parviflorum, T. persistens, T. petiolatum, T. pusillum, T. recurvatum, T. reliquum, T. rivale, T. rugelii, T. sessile, T. simile, T. stamineum, T. sulcatum, T. underwoodii, T. undulatum, T. vaseyi, T. viride, T. viridescens
Subordinate taxa
T. chloropetalum var. chloropetalum, T. chloropetalum var. giganteum
Synonyms T. sessile var. chloropetalum, T. giganteum var. chloropetalum T. sessile var. angustipetalum, T. giganteum var. angustipetalum
Name authority (Torrey) Howell: Fl. N.W. Amer., 661. (1902) (Torrey) J. D. Freeman: Brittonia 27: 55. (1975)
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