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giant trillium, giant white wakerobin, sessile trillium, smallflower trillium, white toadshade

spotted trillium, spotted wakerobin

Rhizomes

horizontal, ± erect, brown, superficially bulblike, short, thick, not brittle.

horizontal, brownish, short, thick, praemorse, not brittle.

Scapes

often several from same rhizome terminus, round in cross section, 2.2–5.8 dm, stout.

1–3, round in cross section, 1.4–4 dm, slender to stout, glabrous.

Bracts

held well above ground, sessile;

blade weakly mottled with scattered, darker green spots, mottling often fading later in season, broadly ovate, 10–20 × 12–15 cm, not glossy, base rounded, apex obtuse.

held well above ground, tips only touching ground in early anthesis, sessile;

blade dark green, mottling darker, mottled to varying degrees, mottling becoming obscure with age, broadly ovate-elliptic to elliptic, 7–15 × 4.6–6.7 cm, not glossy, apex rounded or barely acuminate.

Flower

borne directly upon bracts, erect, fragrance roselike;

sepals conspicuous, spreading, displayed above bracts, pale green, lanceolate, 30–65 × 12–15 mm, margins entire, apex acute;

petals long-lasting, usually erect or slightly spreading, ± connivent, ± concealing stamens and ovary basally, white to creamy white, rarely soft pinkish rose near base, veins prominent but not engraved, not spirally twisted, obovate, more lanceolate in young plants, 4.8–8 × 2.2–3 cm, widest at or just above middle, ± thick-textured, base cuneate, margins entire, ± flat, apex rounded to acute;

stamens erect, greenish white, 15–25 mm;

filaments whitish green, 3–4 mm, slender;

anthers erect, straight, yellow, 11–20 mm, dehiscence latrorse;

connectives straight, extended ca. 1 mm beyond anther sacs, rounded;

ovary green or occasionally purple, ovoid, rounded 6-gonal, 6–11 mm;

stigmas erect to spreading, distinct, sessile, subulate, 4–7 mm, thin distally.

erect, odor faintly spicy and bananalike;

sepals displayed above bracts, spreading almost to horizontal, often suffused or streaked with purple-maroon, lanceolate-linear, 22–50 × 5–7 mm, margins entire, apex rounded-acute, recurving slightly;

petals long-lasting, spreading-erect, ± connivent, ± concealing stamens and ovary or sometimes slightly spreading early, only partially obscuring stamens, becoming more erect and even touching in the manner of Trillium cuneatum when older, clear dark red-maroon, or reddish purple, without muddy or brown tones of many purple sessile species, or, rarely, purple basally and yellow toward apex, or clear sulfur yellow, not spirally twisted, narrowly oblanceolate-spatulate to linear-spatulate, widest above middle, narrowing to apex, 4–7 × 0.7–1.7 cm, thick-textured, margins entire, slightly involute in proximal 1/2, apex acute to round-acute;

stamens erect, 12–20 mm;

filaments dark purple, 2–3 mm, widest at base, much shorter than anther sacs;

anthers erect, straight, brownish purple, 10–16 mm, dehiscence introrse on broad connective;

connectives brown-purple, straight, essentially not extended beyond anther sacs;

ovary dark purple, ovoid, weakly 3-angled to smooth (rarely very obscurely 6-angled), angles often obscured in large, turgid ovary, 8–11 mm;

stigmas erect, divergent-recurved, distinct, purplish, subulate, 2–4 mm, fleshy.

Fruits

green or purplish green, fragrance unknown, ovoid to globose, pulpy, juicy.

dark purplish green, odor not reported, ovoid, obscurely 3- to 6-angled, bearing persistent stigma, 1 × 2 cm, pulpy.

Trillium albidum

Trillium maculatum

Phenology Flowering spring (mid Mar–late Apr). Flowering mid winter–early spring (early Feb–early Apr).
Habitat Moist slopes in rich mixed deciduous-coniferous forests or coniferous stands, brushy thickets on flats, open fields, pastures, and fencerows, dense second-growth coniferous forests, floodplains along streams and larger rivers Rich mesic forests, particularly banks and bluffs of rivers, floodplains, often where quite brushy, rich soils, calcareous soils, alluvium
Elevation 100–200 m (300–700 ft) 0–500 m (0–1600 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; SC
[WildflowerSearch map]
[BONAP county map]
Discussion

In their account of the Carolinas flora, A. E. Radford et al. (1968) did not treat Trillium maculatum or give any explanation of why they omitted this species, which is fairly widely distributed in southeastern South Carolina and morphologically is one of the more distinct sessile trilliums.

Many color forms occur and have been named. In forma luteum J. D. Freeman, for example, all floral organs lack purple pigment, and the petals are clear, soft yellow, whereas forma simulans J. D. Freeman has yellow petals with purple bases, and purple stamens and carpels. These are illustrated in color in F. W. Case and R. B. Case (1997).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 26, p. 106. FNA vol. 26, p. 112.
Parent taxa Liliaceae > Trillium > subg. Phyllantherum Liliaceae > Trillium > subg. Phyllantherum
Sibling taxa
T. angustipetalum, T. catesbaei, T. cernuum, T. chloropetalum, T. cuneatum, T. decipiens, T. decumbens, T. discolor, T. erectum, T. flexipes, T. foetidissimum, T. gracile, T. grandiflorum, T. kurabayashii, T. lancifolium, T. ludovicianum, T. luteum, T. maculatum, T. nivale, T. ovatum, T. parviflorum, T. persistens, T. petiolatum, T. pusillum, T. recurvatum, T. reliquum, T. rivale, T. rugelii, T. sessile, T. simile, T. stamineum, T. sulcatum, T. underwoodii, T. undulatum, T. vaseyi, T. viride, T. viridescens
T. albidum, T. angustipetalum, T. catesbaei, T. cernuum, T. chloropetalum, T. cuneatum, T. decipiens, T. decumbens, T. discolor, T. erectum, T. flexipes, T. foetidissimum, T. gracile, T. grandiflorum, T. kurabayashii, T. lancifolium, T. ludovicianum, T. luteum, T. nivale, T. ovatum, T. parviflorum, T. persistens, T. petiolatum, T. pusillum, T. recurvatum, T. reliquum, T. rivale, T. rugelii, T. sessile, T. simile, T. stamineum, T. sulcatum, T. underwoodii, T. undulatum, T. vaseyi, T. viride, T. viridescens
Name authority J. D. Freeman: Brittonia 27: 48, fig. 11. (1975) Rafinesque: Med. Fl. 2: 103. (1830)
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