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sand clover, thimble clover, tomcat clover, Willdenow's clover

running buffalo clover

Habit Herbs annual, 10–60 cm, glabrous. Herbs perennial, 10–20 cm, glabrous or sparsely pubescent.
Stems

erect, branched.

creeping, branched, rooting at nodes.

Leaves

palmate;

stipules lanceolate or ovate, 1–2 cm, margins dentate or lacerate, apex acuminate;

petiole 1–8 cm;

petiolules 0.5 mm;

leaflets 3, blades linear, lanceolate, or elliptic, 1–5 × 0.2–1.5 cm, base cuneate, veins fine, margins serrate, apex acute, blunt, or retuse, surfaces glabrous.

palmate, opposite or subopposite on erect stems;

stipules lanceolate (on prostrate stems) or ovate (on erect stems), 1–2 cm, margins entire or shallowly serrate, apex acuminate;

petioles 6–15 cm on prostrate stems, 0.5–4 cm on erect stems;

petiolules 1 mm;

leaflets 3, blades broadly ovate or obcordate, 1–2 × 0.5–2.5 cm, base broadly cuneate to rounded, veins fine, margins serrate, apex emarginate or rounded, surfaces glabrous.

Inflorescences

axillary or terminal, 10–50-flowered, globose or ovoid, 1–3.5 × 1–3 cm;

involucres flattened or bowl-shaped, 3–9 mm, when folded, not hiding flowers except proximally, incised 1/4–1/3 their length.

terminal, single or paired at tip of upright stem, 15–30-flowered, globose, 2–3.5 × 2–3.5 cm;

involucres absent.

Peduncles

2–15 cm.

2–7 cm.

Pedicels

erect, 0.5 mm;

bracteoles absent.

elongated and strongly reflexed in fruit, 2–5 mm;

bracteoles lanceolate, 2 mm.

Flowers

12–20 mm;

calyx tubular-campanulate, slit between adaxial lobes, 6–9 mm, glabrous, veins 10–15, tube 3–7 mm, lobes unequal, triangular or subulate, usually 3-fid or shouldered below apex, orifice open;

corolla usually white with purple spot, sometimes lavender or pink or all white, 10–18 mm, banner narrowly oblong, 12–20 × 3–4 mm, apex obtuse or retuse.

9–15 mm;

calyx campanulate, 4–7 mm, glabrous or sparsely pubescent, veins 10, tube 1.3–3.3 mm, lobes subequal, subulate, orifice open;

corolla white, veins often pinkish, 8–14 mm, banner ovate or oblong, 8–14 × 4–5 mm, apex rounded or emarginate.

Legumes

ellipsoid, 3 mm.

oblong, 2.5–3 mm.

Seeds

1 or 2, yellow, mottled, globose-ellipsoid, 1.5–2.5 mm, smooth.

1 or 2, tan to brown, reniform, 1.3–2 mm, smooth.

2n

= 16.

= 16.

Trifolium willdenovii

Trifolium stoloniferum

Phenology Flowering Mar–Jul. Flowering Apr–Jun.
Habitat Roadsides, meadows, lawns, hill­sides, stream margins, open oak chaparral. Grassy openings in upland woods.
Elevation 0–2500 m. (0–8200 ft.) 150–900 m. (500–3000 ft.)
Distribution
from FNA
AZ; CA; ID; OR; WA; BC; Mexico (Baja California)
[WildflowerSearch map]
from FNA
IN; KY; MO; OH; PA; WV
[BONAP county map]
Discussion

Trifolium willdenovii was described by M. Zohary and D. Heller (1984) as one of the most variable clover species in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trifolium stoloniferum was thought to be extinct but was rediscovered in the early 1980s (R. L. Bartgis 1985). It is now known from extant populations scattered throughout much of its original range, in five of the original eight states in which it was found historically (D. J. Crawford et al. 1998); it was discovered in Greene County, Pennsylvania, in 2017, but appears to be extirpated in Arkansas, Illinois, and Kansas. R. J. Hickey et al. (1991) examined genetic diversity of T. stoloniferum populations using allozymes and found low diversity, with some populations appearing genetically uniform. Molecular studies using RAPDs (Crawford et al.) indicated somewhat higher genetic diversity, even in relatively small populations.

Trifolium stoloniferum is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Trifolium Fabaceae > subfam. Faboideae > Trifolium
Sibling taxa
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. wormskioldii
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
Synonyms T. involucratum, T. aciculare, T. nuttallii, T. scabrellum, T. segetum, T. tridentatum, T. tridentatum var. aciculare, T. tridentatum var. scabrellum, T. tridentatum var. segetum, T. tridentatum var. watsonii, T. trimorphum, T. watsonii
Name authority Sprengel: Syst. Veg. 3: 208. (1826) Eaton: Man. Bot. ed. 2, 468. (1818)
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