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woolly clover

running buffalo clover

Habit Herbs annual, 10–20 cm, glabrous or sparsely hairy. Herbs perennial, 10–20 cm, glabrous or sparsely pubescent.
Stems

prostrate, ascending, or erect, branched.

creeping, branched, rooting at nodes.

Leaves

palmate;

stipules ovate or triangular-lanceolate, 0.4–1.2 cm, margins entire, apex acute or acuminate;

petiole 0.5–7 cm;

petiolules to 1 mm;

leaflets 3, blades obovate, obcordate, or elliptic, 0.4–1.5 × 0.3–1 cm, base cuneate, veins moderately prominent, margins denticulate, apex rounded or emarginate, surfaces sparsely hairy abaxially, glabrous adaxially.

palmate, opposite or subopposite on erect stems;

stipules lanceolate (on prostrate stems) or ovate (on erect stems), 1–2 cm, margins entire or shallowly serrate, apex acuminate;

petioles 6–15 cm on prostrate stems, 0.5–4 cm on erect stems;

petiolules 1 mm;

leaflets 3, blades broadly ovate or obcordate, 1–2 × 0.5–2.5 cm, base broadly cuneate to rounded, veins fine, margins serrate, apex emarginate or rounded, surfaces glabrous.

Inflorescences

axillary, 10–20-flowered, subglobose, soon becoming globose, flowers resupinate, calyces densely white-woolly, compacted, inflated, 0.5–1.5 × 0.5–1.5 cm;

involucres a narrow rim, 0.2 mm.

terminal, single or paired at tip of upright stem, 15–30-flowered, globose, 2–3.5 × 2–3.5 cm;

involucres absent.

Peduncles

0.5–1.5 cm.

2–7 cm.

Pedicels

slightly reflexed, to 0.5 mm;

bracteoles cup-shaped, membranous, 0.2 mm.

elongated and strongly reflexed in fruit, 2–5 mm;

bracteoles lanceolate, 2 mm.

Flowers

3–7 mm;

calyx tubular, inflated in fruit, markedly asymmetric-bilabiate, 2–6 mm, woolly adaxially, veins 5–10, connected by lateral veins, tube 1.5–2.5 mm, 4–5 mm in fruit, lobes unequal, subulate, shorter than tube, adaxial spreading or curved, orifice open, abruptly constricted in fruit;

corolla pink, 3–6 mm, banner ovate, 3–6 × 3–6 mm, apex emarginate to crenulate.

9–15 mm;

calyx campanulate, 4–7 mm, glabrous or sparsely pubescent, veins 10, tube 1.3–3.3 mm, lobes subequal, subulate, orifice open;

corolla white, veins often pinkish, 8–14 mm, banner ovate or oblong, 8–14 × 4–5 mm, apex rounded or emarginate.

Legumes

ovoid to globose, 2–3 mm.

oblong, 2.5–3 mm.

Seeds

1 or 2, yellow to brown, mottled, mitten-shaped, 0.9–1.1 mm, smooth, glossy.

1 or 2, tan to brown, reniform, 1.3–2 mm, smooth.

2n

= 16.

= 16.

Trifolium tomentosum

Trifolium stoloniferum

Phenology Flowering Feb–Apr. Flowering Apr–Jun.
Habitat Sandy lawns, fields, meadows, roadsides, clay soils among vernal pools. Grassy openings in upland woods.
Elevation 0–400 m. (0–1300 ft.) 150–900 m. (500–3000 ft.)
Distribution
from FNA
CA; FL; MA; NC; SC; s Europe (Mediterranean); sw Asia; n Africa; Atlantic Islands (Azores) [Introduced in North America; introduced also in South America (Chile), s Africa, Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
IN; KY; MO; OH; PA; WV
[BONAP county map]
Discussion

Trifolium tomentosum is occasionally cultivated as a forage crop (F. J. Hermann 1953) and is becoming weedy in the flora area and in Australia (R. P. Randall 2002).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trifolium stoloniferum was thought to be extinct but was rediscovered in the early 1980s (R. L. Bartgis 1985). It is now known from extant populations scattered throughout much of its original range, in five of the original eight states in which it was found historically (D. J. Crawford et al. 1998); it was discovered in Greene County, Pennsylvania, in 2017, but appears to be extirpated in Arkansas, Illinois, and Kansas. R. J. Hickey et al. (1991) examined genetic diversity of T. stoloniferum populations using allozymes and found low diversity, with some populations appearing genetically uniform. Molecular studies using RAPDs (Crawford et al.) indicated somewhat higher genetic diversity, even in relatively small populations.

Trifolium stoloniferum is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Trifolium Fabaceae > subfam. Faboideae > Trifolium
Sibling taxa
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
Name authority Linnaeus: Sp. Pl. 2: 771. (1753) Eaton: Man. Bot. ed. 2, 468. (1818)
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