Trifolium polyodon |
Trifolium trichocalyx |
|
---|---|---|
Pacific grove clover, woods or Pacific grove clover |
Monterey clover |
|
Habit | Herbs annual, 10–60 cm, glabrous. | Herbs annual, 5–45 cm, sparsely villous to glabrescent. |
Stems | decumbent or ascending, branched. |
prostrate, decumbent, or erect, branched. |
Leaves | palmate; stipules lanceolate to ovate, 0.4–1.8 cm, margins lacerate, apex acute to acuminate; petiole 0.5–6 cm; petiolules to 1 mm; leaflets 3, blades broadly elliptic to broadly obovate, 0.4–2.5 × 0.4–1.5 cm, base cuneate, veins moderately thickened, margins dentate-serrulate, apex rounded or truncate, often retuse, surfaces glabrous. |
palmate; stipules ovate, 0.4–1.2 cm, margins serrate-lacerate, apex acuminate; petiole 1–4 cm; petiolules 0.5 mm; leaflets 3, blades obovate, 0.5–1.5 × 0.4–1 cm, base cuneate, veins moderately prominent, margins denticulate, apex truncate, retuse, or mucronulate, surfaces glabrescent. |
Inflorescences | axillary, 10–25-flowered, subglobose or globose, 1–1.8 × 1–1.8 cm; involucres flattened or bowl-shaped, 4–7 mm, incised 1/2 their length, when folded, not hiding flowers except proximally, lobes 5–12, dentate-lacerate, spinulose. |
axillary or terminal, 2–15-flowered, subglobose, 1.5–1.8 × 0.5–1.5 cm; involucres flattened or vase-shaped, 1–3 mm, when folded, not hiding flowers except proximally, incised 2/3–3/4 their length, lobes 5–15, linear-lanceolate, entire, acuminate. |
Peduncles | 1–2 cm. |
1.5–4 cm. |
Pedicels | straight, to 1 mm; bracteoles absent. |
erect, 1.5 mm; bracteoles absent. |
Flowers | 8–10 mm; calyx campanulate-tubular, not slit between adaxial lobes, 5–7 mm, glabrous, veins 20, tube 1.9–2.4 mm, lobes unequal, 3-fid or laciniate, often appearing 7+-lobed, orifice open; corolla pink or pale purple, 8–9 mm, banner elliptic, 8–9 × 2–3 mm, apex retuse. |
8–11 mm; calyx campanulate-tubular, not slit between adaxial lobes, 6–9 mm, usually densely pubescent, rarely sparsely so, veins 10, tube 2.5–4.5 mm, lobes unequal, lanceolate-subulate, orifice open; corolla light purple with darker purple keel petals, 6–10 mm, banner narrowly obovate, 4–5 × 1 mm, apex retuse. |
Legumes | ellipsoid, 3.5–4 mm. |
sessile, oblong, 5 mm. |
Seeds | 2, dark brown, mottled, ovoid or reniform, 1.7–1.9 mm, smooth. |
3–6(–9), pale brown, mottled purple, globose to mitten-shaped, 0.8–1 mm, smooth to slightly roughened, semiglossy. |
Trifolium polyodon |
Trifolium trichocalyx |
|
Phenology | Flowering Apr–Jun. | Flowering Apr–May. |
Habitat | Along streams, moist meadows. | Sandy, rich soils in open Monterey pine forests, often after fire. |
Elevation | 0–150 m. (0–500 ft.) | 0–50 m. (0–200 ft.) |
Distribution |
CA |
CA |
Discussion | The taxonomic status of Trifolium polyodon, known only from Monterey County, has long been debated. W. L. Jepson (1936) considered it a variety of T. tridentatum Lindley (= T. willdenovii). The first to consider it as a variety of T. variegatum was J. S. Martin (1943), who never formally published a new combination; an invalid combination was made by M. Zohary and D. Heller (1984). D. Isely (1998) included it in his interpretation of T. variegatum as phase 5 of that species; he speculated that T. polyodon might have originated as a hybrid of T. variegatum and T. willdenovii. Molecular studies (N. W. Ellison et al. 2006) showed that T. polyodon is closely related to T. variegatum but is distinct; it also appears to be related to T. cyathiferum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Trifolium trichocalyx is an extremely rare species known only from Mendocino and Monterey counties. D. I. Axelrod (1982) suggested that T. trichocalyx might have originated as a hybrid between T. microcephalum and T. variegatum, but this is not supported by molecular studies (N. W. Ellison et al. 2006). It appears to be fire-adapted and appeared in large numbers shortly after a fire in 1987 (U.S. Fish and Wildlife Service 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | T. tridentatum var. polyodon | T. oliganthum var. trichocalyx |
Name authority | Greene: Pittonia 3: 215. (1897) | A. Heller: Muhlenbergia 1: 55. (1904) |
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