Trifolium macrocephalum |
Trifolium cyathiferum |
|
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big-head clover, large-head clover |
bowl clover, cup clover |
|
Habit | Herbs perennial, 5–25 cm, villous. | Herbs annual, 5–50 cm, glabrous. |
Stems | erect or ascending, branched. |
erect, branched. |
Leaves | palmate; stipules ovate, obovate, or oblong, 1–3 cm, margins entire, irregularly lobed, or serrate, apex acute or acuminate; petiole 1–14 cm; petiolules 0.9–1.2 mm; leaflets (5–)7–9, blades broadly to narrowly obovate, often folded, 1–2.7 × 0.4–1.1 cm, base cuneate, veins thickened, especially distally, margins serrulate, apex rounded or truncate, apiculate, surfaces villous abaxially, sparsely villous to glabrate adaxially. |
palmate; stipules ovate to lanceolate, 0.6–1.2 cm, margins entire, toothed, or lacerate, apex acute or aristate; petiole 1–10 cm; petiolules to 0.5 mm; leaflets 3, blades obovate, elliptic, or oblanceolate, 0.5–2.5 × 0.3–1.5 cm, base cuneate, veins prominent, green, margins finely toothed, apex rounded, blunt, or retuse, surfaces glabrous. |
Inflorescences | terminal, 20–32-flowered, globose or ovoid-ellipsoid, 2.5–8 × 3–7 cm; involucres absent. |
axillary or terminal, 1–35-flowered, subglobose or ovoid, 0.5–1.8 × 0.5–2 cm; involucres broadly bowl-shaped, 4–22 mm, shallowly incised, lobes 3–15, toothed, broad, acute. |
Peduncles | 2–8 cm. |
1–8 cm. |
Pedicels | erect, 1–1.5 mm; bracteoles broadly ovate, membranous, to 0.5 mm, truncate. |
straight, 0.1–0.2 mm; bracteoles absent. |
Flowers | 20–30 mm; calyx campanulate, 10–22 mm, villous, veins 10–15, tube 2.5–4 mm, lobes subequal, subulate, plumose, orifice open; corolla white, creamy white, or pinkish, keel petals deep pink, 20–28 mm, banner ovate or oblong, 20–28 × 10–13 mm, apex rounded or slightly emarginate. |
7–15 mm; calyx campanulate, inflated in fruit, 6–13 mm, glabrous, veins 13–20, tube 3–7 mm, lobes unequal, strongly oblique, broadly triangular, apex setaceous, adaxial unbranched, abaxial and lateral conspicuously 2- or 3-fid, segments glabrous, sometimes flattened, orifice open; corolla usually creamy white or rose to pink, sometimes whitish with pinkish tips, 6–13 mm, banner obovate or elliptic, proximally inflated in fruit, distally narrowed into twisted tip, 6–13 × 3–6 mm, apex rounded to broadly acute. |
Legumes | ovoid, 4–5 mm. |
ellipsoid, 2.5–3 mm. |
Seeds | 1 or 2, yellow or reddish, mitten-shaped to ellipsoid, 2.5–3 mm, smooth. |
1 or 2, yellow-brown, ovoid, 0.4–0.6 mm, smooth. |
2n | = 32, 48. |
= 16. |
Trifolium macrocephalum |
Trifolium cyathiferum |
|
Phenology | Flowering Apr–Jun. | Flowering Apr–Oct. |
Habitat | Rocky places, hard, compacted clay-gumbo, lava beds, sage-covered slopes, full sun. | Wet meadows, roadsides, fields. |
Elevation | 80–2500 m. [260–8200 ft.] | 0–2700 m. [0–8900 ft.] |
Distribution |
CA; ID; NV; OR; WA
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AK; CA; ID; MT; NV; OR; WA; BC; YT [Introduced in Asia (China, Japan)]
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Discussion | Trifolium macrocephalum has the largest inflorescences of any clover. Trifolium megacephalum Nuttall (1818) is an illegitimate replacement name for Lupinaster macrocephalum Pursh. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Trifolium cyathiferum may be of hybrid origin; phylogenetic studies indicate it has the same trnL intron sequence as T. buckwestiorum, T. polyodon, and T. variegatum and nrDNA and combined analyses place it in a clade with other species (T. barbigerum, T. fucatum, T. jokerstii, and T. physanthum Hooker & Arnott) that have inflated fruiting corollas (N. W. Ellison et al. 2006). The single record of Trifolium cyathiferum labeled as from Utah is likely mislabeled (S. L. Welsh et al. 2008). Populations in Yukon may be introductions. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Lupinaster macrocephalum | |
Name authority | (Pursh) Poiret in J. Lamarck et al.: Encycl., suppl. 5: 336. (1817) | Lindley: Bot. Reg. 13: sub plate 1070. (1827) |
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