Trifolium macrocephalum |
Trifolium columbinum |
|
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big-head clover, large-head clover |
olive clover |
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Habit | Herbs perennial, 5–25 cm, villous. | Herbs annual, 8–30 cm, canescent. |
Stems | erect or ascending, branched. |
erect, branched from base and distally, or unbranched. |
Leaves | palmate; stipules ovate, obovate, or oblong, 1–3 cm, margins entire, irregularly lobed, or serrate, apex acute or acuminate; petiole 1–14 cm; petiolules 0.9–1.2 mm; leaflets (5–)7–9, blades broadly to narrowly obovate, often folded, 1–2.7 × 0.4–1.1 cm, base cuneate, veins thickened, especially distally, margins serrulate, apex rounded or truncate, apiculate, surfaces villous abaxially, sparsely villous to glabrate adaxially. |
palmate; stipules ovate, 0.4–1.2 cm, margins entire, ciliate, apex often purple, acuminate, setaceous; petiole 0.5–8 cm; petiolules 0.5–1 mm; leaflets 3, blades obovate or elliptic, 0.5–3 × 0.3–1.2 cm, base cuneate, veins obscure or slightly thickened, margins dentate, apex rounded or emarginate, surfaces pilose. |
Inflorescences | terminal, 20–32-flowered, globose or ovoid-ellipsoid, 2.5–8 × 3–7 cm; involucres absent. |
terminal or axillary, 25–50-flowered, ovoid-ellipsoid, 1–2.5 × 1.2–2.5 cm; involucres absent. |
Peduncles | 2–8 cm. |
3–11 cm. |
Pedicels | erect, 1–1.5 mm; bracteoles broadly ovate, membranous, to 0.5 mm, truncate. |
absent; bracteoles cuplike, to 0.5 mm. |
Flowers | 20–30 mm; calyx campanulate, 10–22 mm, villous, veins 10–15, tube 2.5–4 mm, lobes subequal, subulate, plumose, orifice open; corolla white, creamy white, or pinkish, keel petals deep pink, 20–28 mm, banner ovate or oblong, 20–28 × 10–13 mm, apex rounded or slightly emarginate. |
6–10 mm; calyx tubular, 8–14 mm, densely pubescent, veins 20–30, tube 1–2 mm, lobes nearly equal, appearing rigidly erect, linear-setaceous, plumose, sinuses acute, orifice open; corolla sometimes absent, red-violet, often with white tips, 4–6 mm, banner ovate-oblong, 7–8 × 2–3 mm, apex broadly rounded, apiculate. |
Legumes | ovoid, 4–5 mm. |
obovoid, 3–3.5 mm. |
Seeds | 1 or 2, yellow or reddish, mitten-shaped to ellipsoid, 2.5–3 mm, smooth. |
1, yellow-brown, ellipsoid, 2–2.5 mm, smooth. |
2n | = 32, 48. |
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Trifolium macrocephalum |
Trifolium columbinum |
|
Phenology | Flowering Apr–Jun. | Flowering Apr–May. |
Habitat | Rocky places, hard, compacted clay-gumbo, lava beds, sage-covered slopes, full sun. | Grassy slopes, meadows, heavy adobe/clay soils, oak woodland foothills. |
Elevation | 80–2500 m. (300–8200 ft.) | 0–1400 m. (0–4600 ft.) |
Distribution |
CA; ID; NV; OR; WA
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CA |
Discussion | Trifolium macrocephalum has the largest inflorescences of any clover. Trifolium megacephalum Nuttall (1818) is an illegitimate replacement name for Lupinaster macrocephalum Pursh. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
D. Isely (1998) stated that Central Valley populations of Trifolium columbinum appeared to be extinct and populations outside that region (and here recognized as the same taxon) were similar in calyx and corolla proportions. It is rather widespread in California, ranging from Humboldt County southward to Santa Barbara County, and eastward into counties ranging from Shasta southward to Inyo, Kern, and Placer counties. Trifolium albopurpureum var. olivaceum (Greene) Isely and T. columbinum var. olivaceum (Greene) Jepson are illegitimate names that pertain here (D. J. Keil 2019). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | Fabaceae > subfam. Faboideae > Trifolium | Fabaceae > subfam. Faboideae > Trifolium |
Sibling taxa | ||
Synonyms | Lupinaster macrocephalum | T. albopurpureum var. columbinum, T. columbinum var. argillorum, T. olivaceum, T. olivaceum var. columbinum, T. olivaceum var. griseum |
Name authority | (Pursh) Poiret in J. Lamarck et al.: Encycl., suppl. 5: 336. (1817) | Greene: Pittonia 1: 4. (1887) |
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