The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

long-stalk clover

olive clover

Habit Herbs perennial, 5–46 cm, slightly pilose. Herbs annual, 8–30 cm, canescent.
Stems

erect or ascending, cespitose, unbranched to sparsely branched distally.

erect, branched from base and distally, or unbranched.

Leaves

palmate;

stipules ovate to lanceolate, 1–5 cm, margins usually entire, sometimes slightly serrate proximally, apex acute or acuminate;

petiole 1–13 cm;

petiolules to 1 mm;

leaflets 3, blades linear, lanceolate to elliptic, or oblong, cauline sometimes ovate, 0.8–6 × 0.2–1.6 cm, base cuneate, veins slightly to very thickened, arching, margins entire or serrate, apex acute, surfaces glabrous abaxially, usually appressed-pubescent adaxially.

palmate;

stipules ovate, 0.4–1.2 cm, margins entire, ciliate, apex often purple, acuminate, setaceous;

petiole 0.5–8 cm;

petiolules 0.5–1 mm;

leaflets 3, blades obovate or elliptic, 0.5–3 × 0.3–1.2 cm, base cuneate, veins obscure or slightly thickened, margins dentate, apex rounded or emarginate, surfaces pilose.

Inflorescences

terminal, 10–40-flowered, globose, ellipsoid, obconic, or subglobose, 1–3.5 × 1–3.2 cm;

involucres absent.

terminal or axillary, 25–50-flowered, ovoid-ellipsoid, 1–2.5 × 1.2–2.5 cm;

involucres absent.

Peduncles

1–30 cm.

3–11 cm.

Pedicels

erect or reflexed in fruit, 0.5–2 mm;

bracteoles scalelike, 0.5 mm.

absent;

bracteoles cuplike, to 0.5 mm.

Flowers

10–19 mm;

calyx tubular, 4–10 mm, hairy or glabrous, veins 5–10, tube 2.5–3 mm, lobes unequal, linear, shorter than or to 6 times tube, orifice open;

corolla white, cream, pink, lavender, or purple, sometimes pale with purple tips, 10–18 mm, petals not beaked except subsp. multipedunculatum, banner ovate to oblanceolate, 10–18 × 3–4 mm, apex usually not beaked, usually obtuse, acute, or acuminate, rarely emarginate;

ovaries pubescent distally.

6–10 mm;

calyx tubular, 8–14 mm, densely pubescent, veins 20–30, tube 1–2 mm, lobes nearly equal, appearing rigidly erect, linear-setaceous, plumose, sinuses acute, orifice open;

corolla sometimes absent, red-violet, often with white tips, 4–6 mm, banner ovate-oblong, 7–8 × 2–3 mm, apex broadly rounded, apiculate.

Legumes

oblong to ellipsoid, 4–5 mm.

obovoid, 3–3.5 mm.

Seeds

1–4, dark brown, flattened globose, 2–3 mm, smooth.

1, yellow-brown, ellipsoid, 2–2.5 mm, smooth.

Trifolium longipes

Trifolium columbinum

Phenology Flowering Apr–May.
Habitat Grassy slopes, meadows, heavy adobe/clay soils, oak woodland foothills.
Elevation 0–1400 m. [0–4600 ft.]
Distribution
from USDA
w United States
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
Discussion

Subspecies 12 (12 in the flora).

Trifolium longipes encompasses a morphologically diverse and taxonomically difficult suite of populations. J. M. Gillett (1969) united T. longipes with T. rusbyi and differentiated ten subspecies, with T. neurophyllum recognized as distinct. M. Zohary and D. Heller (1984) maintained T. rusbyi (with seven subspecies, including subsp. neurophyllum) as distinct from T. longipes (with five subspecies). Analyses of molecular data support the union of T. longipes and T. rusbyi (N. W. Ellison et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. Isely (1998) stated that Central Valley populations of Trifolium columbinum appeared to be extinct and populations outside that region (and here recognized as the same taxon) were similar in calyx and corolla proportions. It is rather widespread in California, ranging from Humboldt County southward to Santa Barbara County, and eastward into counties ranging from Shasta southward to Inyo, Kern, and Placer counties.

Trifolium albopurpureum var. olivaceum (Greene) Isely and T. columbinum var. olivaceum (Greene) Jepson are illegitimate names that pertain here (D. J. Keil 2019).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Banner, wing, and, often, keel petals beaked.
subsp. multipedunculatum
1. Banner, wing, and keel petals without beaks, banner apex acute to attenuate.
→ 2
2. Pedicels erect or proximalmost horizontal or slightly reflexed in fruit.
→ 3
3. Herbs rhizomatous; roots not fusiform, sometimes thickened.
→ 4
4. Calyx lobes lax, usually slightly longer than tube length, tube veins not promi­nent; leaflet blades lanceolate, elliptic to oblong, or ovate.
subsp. hansenii
4. Calyx lobes often ± rigid, 6 times tube length, tube veins usually prominent; leaflet blades linear to elliptic-linear or broadly elliptic.
subsp. elmeri
3. Herbs not rhizomatous (stems ascending from crowns); roots thickened, often fusiform.
→ 5
5. Banner and wing apices slender-acuminate.
subsp. longipes
5. Banner and wing apices usually acute or obtuse, rarely emarginate.
→ 6
6. Calyces and distal ends of peduncles densely pilose; banner apex acute.
subsp. atrorubens
6. Calyces and distal ends of peduncles sparsely pilose or glabrous; banner apex usually broad, acute or obtuse, rarely emarginate.
subsp. pedunculatum
2. Pedicels (at least proximal ones, usually all), strongly reflexed in fruit.
→ 7
7. Herbs with thickened roots, often fusiform, with crown of buried stems.
subsp. pygmaeum
7. Herbs usually rhizomatous, if rarely with roots ± thickened, then with crown of rhizomes above thickened portion, or with slender taproots bearing a crown of wiry, pilose stems.
→ 8
8. Pedicels slender, 1.5–2 mm; herbs with slender taproots, bearing crown of wiry, pilose stems.
subsp. oreganum
8. Pedicels stout, to 1 mm; herbs rhizomatous.
→ 9
9. Inflorescences globose or ellipsoid; proximal pedicels more strongly reflexed than distal ones.
→ 10
10. Inflorescences 18–22-flowered, globose, 2.2–2.5 cm diam.; flow­ers 10–13 mm, corollas purple, sometimes tips white.
subsp. shastense
10. Inflorescences 28–32-flowered, ellipsoid, 3–3.2 cm diam.; flow­ers 14–16 mm, corollas lavender.
subsp. caurinum
9. Inflorescences hemispheric, obconic, or sub­glo­bose; all pedicels strongly reflexed.
→ 11
11. Calyces sparsely pilose or curly-pilose at sinuses, often glabrous proximally; corollas white, cream, or violet.
subsp. reflexum
11. Calyces white-villous, veins and lobes green; corollas purple-violet or lavender.
subsp. neurophyllum
Source FNA vol. 11. FNA vol. 11. Treatment authors: Michael A. Vincent, John M. Gillett†.
Parent taxa Fabaceae > subfam. Faboideae > Trifolium Fabaceae > subfam. Faboideae > Trifolium
Sibling taxa
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
Subordinate taxa
T. longipes subsp. atrorubens, T. longipes subsp. caurinum, T. longipes subsp. elmeri, T. longipes subsp. hansenii, T. longipes subsp. longipes, T. longipes subsp. multipedunculatum, T. longipes subsp. neurophyllum, T. longipes subsp. oreganum, T. longipes subsp. pedunculatum, T. longipes subsp. pygmaeum, T. longipes subsp. reflexum, T. longipes subsp. shastense
Synonyms T. albopurpureum var. columbinum, T. columbinum var. argillorum, T. olivaceum, T. olivaceum var. columbinum, T. olivaceum var. griseum
Name authority Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 314. (1838) Greene: Pittonia 1: 4. (1887)
Web links