Trifolium eriocephalum |
Trifolium stoloniferum |
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woolly-head clover |
running buffalo clover |
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Habit | Herbs perennial, 12–46 cm, sparsely hairy to densely villous, appressed-pubescent, or glabrate. | Herbs perennial, 10–20 cm, glabrous or sparsely pubescent. | ||||||||||||||||||||
Stems | erect or decumbent, branched. |
creeping, branched, rooting at nodes. |
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Leaves | palmate; stipules lanceolate or ovate, 1–5 cm, margins entire or denticulate, apex acuminate or acute; petiole 1–15 cm; petiolules 1 mm; leaflets 3, blades linear, linear-elliptic, elliptic, lanceolate, lanceolate-oblong, or obovate, 1–5 × 0.4–1.2 cm, base cuneate, veins fine or ± thickened, margins denticulate or serrulate, apex acuminate or acute, surfaces appressed-pubescent abaxially, glabrous, densely villous, or sparsely or densely pilose adaxially. |
palmate, opposite or subopposite on erect stems; stipules lanceolate (on prostrate stems) or ovate (on erect stems), 1–2 cm, margins entire or shallowly serrate, apex acuminate; petioles 6–15 cm on prostrate stems, 0.5–4 cm on erect stems; petiolules 1 mm; leaflets 3, blades broadly ovate or obcordate, 1–2 × 0.5–2.5 cm, base broadly cuneate to rounded, veins fine, margins serrate, apex emarginate or rounded, surfaces glabrous. |
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Inflorescences | terminal or axillary, 25–70-flowered, inverted or horizontal, ellipsoid or globose to subglobose, 2–3 × 1–3 cm, rachis prolonged beyond flowers; involucres absent. |
terminal, single or paired at tip of upright stem, 15–30-flowered, globose, 2–3.5 × 2–3.5 cm; involucres absent. |
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Peduncles | bent distally, proximal to flowers, 3–18 cm. |
2–7 cm. |
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Pedicels | reflexed, to 0.3 mm; bracteoles absent. |
elongated and strongly reflexed in fruit, 2–5 mm; bracteoles lanceolate, 2 mm. |
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Flowers | 9–18 mm; calyx campanulate, curved in fruit, 4–11 mm, villous, veins 10, tube 1.5–2.5 mm, lobes unequal, 2–3 times tube, subulate, often strongly curved and twisted, plumose, orifice open; corolla white, cream, or purple, 8–16 mm, banner oblanceolate, 8–16 × 3–5 mm, apex rounded or retuse, recurved; ovaries pubescent distally. |
9–15 mm; calyx campanulate, 4–7 mm, glabrous or sparsely pubescent, veins 10, tube 1.3–3.3 mm, lobes subequal, subulate, orifice open; corolla white, veins often pinkish, 8–14 mm, banner ovate or oblong, 8–14 × 4–5 mm, apex rounded or emarginate. |
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Legumes | ovoid, 2–3.5 mm. |
oblong, 2.5–3 mm. |
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Seeds | (1 or)2–4, yellow-brown to brown, mitten-shaped or subglobose, 1.2–2 mm, smooth. |
1 or 2, tan to brown, reniform, 1.3–2 mm, smooth. |
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2n | = 16. |
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Trifolium eriocephalum |
Trifolium stoloniferum |
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Phenology | Flowering Apr–Jun. | |||||||||||||||||||||
Habitat | Grassy openings in upland woods. | |||||||||||||||||||||
Elevation | 150–900 m. [500–3000 ft.] | |||||||||||||||||||||
Distribution |
w United States
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IN; KY; MO; OH; PA; WV |
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Discussion | Subspecies 6 (6 in the flora). Trifolium eriocephalum was revised by J. M. Gillett (1971). The subspecies exhibit unusually complex distributional patterns, and overlapping morphological features sometimes make identifications problematic (M. Zohary and D. Heller 1984). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Trifolium stoloniferum was thought to be extinct but was rediscovered in the early 1980s (R. L. Bartgis 1985). It is now known from extant populations scattered throughout much of its original range, in five of the original eight states in which it was found historically (D. J. Crawford et al. 1998); it was discovered in Greene County, Pennsylvania, in 2017, but appears to be extirpated in Arkansas, Illinois, and Kansas. R. J. Hickey et al. (1991) examined genetic diversity of T. stoloniferum populations using allozymes and found low diversity, with some populations appearing genetically uniform. Molecular studies using RAPDs (Crawford et al.) indicated somewhat higher genetic diversity, even in relatively small populations. Trifolium stoloniferum is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 11. | FNA vol. 11. | ||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||
Name authority | Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 313. (1838) | Eaton: Man. Bot. ed. 2, 468. (1818) | ||||||||||||||||||||
Web links |