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woolly-head clover

Persian clover, reversed clover

Habit Herbs perennial, 12–46 cm, sparsely hairy to densely villous, appressed-pubescent, or glabrate. Herbs annual, 20–60 cm, glabrous or glabrescent.
Stems

erect or decumbent, branched.

ascending, erect, or procumbent, branched.

Leaves

palmate;

stipules lanceolate or ovate, 1–5 cm, margins entire or denticulate, apex acuminate or acute;

petiole 1–15 cm;

petiolules 1 mm;

leaflets 3, blades linear, linear-elliptic, elliptic, lanceolate, lanceolate-oblong, or obovate, 1–5 × 0.4–1.2 cm, base cuneate, veins fine or ± thickened, margins denticulate or serrulate, apex acuminate or acute, surfaces appressed-pubescent abaxially, glabrous, densely villous, or sparsely or densely pilose adaxially.

palmate;

stipules lanceolate or ovate, 0.5–1.8 cm, margins entire, apex acuminate-filiform;

petiole 0.5–20 cm;

petiolules to 1 mm;

leaflets 3, blades obovate, ovate, elliptic, or rhombic, 1–3 × 0.5–3 cm, base cuneate, veins moderately prominent, margins spinulose-dentate, apex rounded or broadly acute, surfaces hairy abaxially, glabrous adaxially.

Inflorescences

terminal or axillary, 25–70-flowered, inverted or horizontal, ellipsoid or globose to subglobose, 2–3 × 1–3 cm, rachis prolonged beyond flowers;

involucres absent.

axillary, 15–30-flowered, subglobose to globose, flowers resupinate, forming stellate-spreading clusters in fruit, 0.5–1.6 × 0.8–1.5 cm;

involucres a narrow rim, 0.2 mm.

Peduncles

bent distally, proximal to flowers, 3–18 cm.

1–6 cm.

Pedicels

reflexed, to 0.3 mm;

bracteoles absent.

slightly reflexed, to 0.5 mm;

bracteoles minute or absent.

Flowers

9–18 mm;

calyx campanulate, curved in fruit, 4–11 mm, villous, veins 10, tube 1.5–2.5 mm, lobes unequal, 2–3 times tube, subulate, often strongly curved and twisted, plumose, orifice open;

corolla white, cream, or purple, 8–16 mm, banner oblanceolate, 8–16 × 3–5 mm, apex rounded or retuse, recurved;

ovaries pubescent distally.

5–6 mm;

calyx tubular-campanulate, inflated in fruit, markedly asymmetric-bilabiate, slit between adaxial lobes, 2.5–3 mm, 8–11 mm in fruit, hairy, short-hairy or glabrescent in fruit, veins 10, connected by lateral veins, tube 2 mm, 7–9 mm in fruit, lobes green, erect, unequal, triangular, divergent, linear-lanceolate, unequal in fruit, orifice open;

corolla pink to purple, 5–8 mm, banner oblong, 5–8 × 2 mm, apex emarginate to crenulate.

Legumes

ovoid, 2–3.5 mm.

lenticular, 1.8–2.2 mm.

Seeds

(1 or)2–4, yellow-brown to brown, mitten-shaped or subglobose, 1.2–2 mm, smooth.

1, ovoid, dark purple, olive green, yellow, or reddish brown, 1.5–1.9 mm, smooth, dull.

2n

= 14, 16, 32.

Trifolium eriocephalum

Trifolium resupinatum

Phenology Flowering Mar–May.
Habitat Wet meadows, lawns, road­sides, fields, waste places.
Elevation 0–700 m. [0–2300 ft.]
Distribution
from USDA
w United States
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; DC; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MO; MS; NC; ND; OH; OK; OR; PA; SC; SD; TN; TX; VA; VT; WA; WI; WV; ON; c Europe; s Europe; sw Asia; n Africa; intro­duced also in s South America (Argentina, Uruguay); s Africa; Pacific Islands (Hawaii, New Zealand); Australia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 6 (6 in the flora).

Trifolium eriocephalum was revised by J. M. Gillett (1971). The subspecies exhibit unusually complex distributional patterns, and overlapping morphological features sometimes make identifications problematic (M. Zohary and D. Heller 1984).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Widespread use of Trifolium resupinatum as a forage crop began in the late 1920s, after it began to flourish in Louisiana following a flood of the Mississippi River (E. A. Hollowell 1943). It is used in the southern United States as a pasture plant and for production of hay, and has spread widely from cultivation.

Reports of Trifolium resupinatum for New Brunswick and Quebec are based on old collections; since the species has not been recollected for many years in either province, they are excluded here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ovules usually 2, rarely 1 or 3; calyx lobes usually green, rarely purple, straight, subequal, 3–4 timestube.
→ 2
2. Herbs densely villous; leaflet blades of basal and median leaves elliptic or lanceolate-oblong; roots cylindric, thickened.
subsp. eriocephalum
2. Herbs slightly villous, appressed-pubescent, or glabrate; leaflet blades of basal and med­ian leaves usually narrowly lanceolate or lin­ear, rarely narrowly elliptic; roots fusiform-tuberous.
subsp. cascadense
1. Ovules 3 or 4; calyx lobes usually purple, contorted, abaxial lobes longer, lateral lobes 1–1.5 times tube.
→ 3
3. Leaflet blades of basal leaves linear or linear-elliptic, apex acute, surfaces sparsely or densely hairy adaxially; roots cylindric, thickened, branched, often rhizomelike.
→ 4
4. Blades of leaflets of median leaves linear or linear-elliptic; herbs slightly villous.
subsp. cusickii
4. Blades of leaflets of median leaves broadly elliptic; herbs densely villous.
subsp. villiferum
3. Leaflet blades of basal leaves obovate, apex rounded to retuse, surfaces usually glabrous adaxially, sometimes sparsely hairy; roots fusiform-tuberous.
→ 5
5. Inflorescences 2.5 cm diam., significantly overtopping distal leaves; leaflet blades of median leaves broadly lanceolate, surfaces sparsely hairy or glabrous adaxially.
subsp. arcuatum
5. Inflorescences 1 cm diam., scarcely over­topping distal leaves; leaflet blades of median leaves narrowly lanceolate or linear, surfaces densely villous adaxially.
subsp. martinii
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Trifolium Fabaceae > subfam. Faboideae > Trifolium
Sibling taxa
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
Subordinate taxa
T. eriocephalum subsp. arcuatum, T. eriocephalum subsp. cascadense, T. eriocephalum subsp. cusickii, T. eriocephalum subsp. eriocephalum, T. eriocephalum subsp. martinii, T. eriocephalum subsp. villiferum
Name authority Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 313. (1838) Linnaeus: Sp. Pl. 2: 771. (1753)
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