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woolly-head clover

andrews' clover, Gray's clover

Habit Herbs perennial, 12–46 cm, sparsely hairy to densely villous, appressed-pubescent, or glabrate. Herbs annual, 10–40 cm, densely pubescent or glabrate.
Stems

erect or decumbent, branched.

erect, branched.

Leaves

palmate;

stipules lanceolate or ovate, 1–5 cm, margins entire or denticulate, apex acuminate or acute;

petiole 1–15 cm;

petiolules 1 mm;

leaflets 3, blades linear, linear-elliptic, elliptic, lanceolate, lanceolate-oblong, or obovate, 1–5 × 0.4–1.2 cm, base cuneate, veins fine or ± thickened, margins denticulate or serrulate, apex acuminate or acute, surfaces appressed-pubescent abaxially, glabrous, densely villous, or sparsely or densely pilose adaxially.

palmate;

stipules ovate, 0.5–1.5 cm, margins toothed or lacerate, apex acute-acuminate;

petiole 1–15 cm;

petiolules 1 mm;

leaflets 3, blades elliptic, oblanceolate, or obovate, 2–2.5 × 1–1.5 cm, base cuneate, veins fine, margins dentate-serrate to crenulate, teeth shortly aristate, apex obtuse or broadly acute, surfaces pubescent or glabrate.

Inflorescences

terminal or axillary, 25–70-flowered, inverted or horizontal, ellipsoid or globose to subglobose, 2–3 × 1–3 cm, rachis prolonged beyond flowers;

involucres absent.

axillary or terminal, 5–30-flowered, subglobose or globose, 1.8–3 × 2–3 cm;

involucres bowl-shaped, 10–25 mm, lobes 6–16, sharply setaceous-toothed, sinuses shallow.

Peduncles

bent distally, proximal to flowers, 3–18 cm.

2–15 cm.

Pedicels

reflexed, to 0.3 mm;

bracteoles absent.

straight, 0.5 mm;

bracteoles absent.

Flowers

9–18 mm;

calyx campanulate, curved in fruit, 4–11 mm, villous, veins 10, tube 1.5–2.5 mm, lobes unequal, 2–3 times tube, subulate, often strongly curved and twisted, plumose, orifice open;

corolla white, cream, or purple, 8–16 mm, banner oblanceolate, 8–16 × 3–5 mm, apex rounded or retuse, recurved;

ovaries pubescent distally.

10–17 mm;

calyx campanulate, 6–13 mm, pubescent, veins 5, tube 3–5 mm, lobes unequal, setaceous, abaxial inconspicuously 2- or 3-fid, adaxial unbranched, segments plumose, orifice open;

corolla usually lavender or purple, sometimes purple with white tips, 8–16 mm, banner oblong, proximally inflated in fruit, distally narrowed into twisted tip, 3–5 × 10–13 mm, apex obtuse, truncate, or emarginate.

Legumes

ovoid, 2–3.5 mm.

stipitate, ovoid-ellipsoid, 2.5–4 mm.

Seeds

(1 or)2–4, yellow-brown to brown, mitten-shaped or subglobose, 1.2–2 mm, smooth.

1 or 2, pale brown, mottled, ellipsoid to mitten-shaped, 1.6–2 mm, rugose.

Trifolium eriocephalum

Trifolium grayi

Phenology Flowering Apr–Jun.
Habitat Wet meadows, foothill slopes, pine woodlands.
Elevation 0–600 m. [0–2000 ft.]
Distribution
from USDA
w United States
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Subspecies 6 (6 in the flora).

Trifolium eriocephalum was revised by J. M. Gillett (1971). The subspecies exhibit unusually complex distributional patterns, and overlapping morphological features sometimes make identifications problematic (M. Zohary and D. Heller 1984).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trifolium grayi, which ranges from San Luis Obispo County in the south to Mendocino County in the north, and eastward into Amador, Sacramento, and Tuolumne counties, has long been considered a variety of T. barbigerum and some authors claim that intermediates between the two species are encountered (J. S. Martin 1943; D. Isely 1998); others state that the two taxa are distinct (L. F. McDermott 1910; M. A Vincent and R. Morgan 1998).

Trifolium andrewsii (A. Gray) A. Heller is an illegitimate superfluous name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Fabaceae > subfam. Faboideae > Trifolium Fabaceae > subfam. Faboideae > Trifolium
Sibling taxa
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
Subordinate taxa
T. eriocephalum subsp. arcuatum, T. eriocephalum subsp. cascadense, T. eriocephalum subsp. cusickii, T. eriocephalum subsp. eriocephalum, T. eriocephalum subsp. martinii, T. eriocephalum subsp. villiferum
Key
1. Ovules usually 2, rarely 1 or 3; calyx lobes usually green, rarely purple, straight, subequal, 3–4 timestube.
→ 2
2. Herbs densely villous; leaflet blades of basal and median leaves elliptic or lanceolate-oblong; roots cylindric, thickened.
subsp. eriocephalum
2. Herbs slightly villous, appressed-pubescent, or glabrate; leaflet blades of basal and med­ian leaves usually narrowly lanceolate or lin­ear, rarely narrowly elliptic; roots fusiform-tuberous.
subsp. cascadense
1. Ovules 3 or 4; calyx lobes usually purple, contorted, abaxial lobes longer, lateral lobes 1–1.5 times tube.
→ 3
3. Leaflet blades of basal leaves linear or linear-elliptic, apex acute, surfaces sparsely or densely hairy adaxially; roots cylindric, thickened, branched, often rhizomelike.
→ 4
4. Blades of leaflets of median leaves linear or linear-elliptic; herbs slightly villous.
subsp. cusickii
4. Blades of leaflets of median leaves broadly elliptic; herbs densely villous.
subsp. villiferum
3. Leaflet blades of basal leaves obovate, apex rounded to retuse, surfaces usually glabrous adaxially, sometimes sparsely hairy; roots fusiform-tuberous.
→ 5
5. Inflorescences 2.5 cm diam., significantly overtopping distal leaves; leaflet blades of median leaves broadly lanceolate, surfaces sparsely hairy or glabrous adaxially.
subsp. arcuatum
5. Inflorescences 1 cm diam., scarcely over­topping distal leaves; leaflet blades of median leaves narrowly lanceolate or linear, surfaces densely villous adaxially.
subsp. martinii
Synonyms T. barbigerum var. andrewsii, T. barbigerum var. lilacinum, T. lilacinum
Name authority Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 313. (1838) Lojacono: Nuovo Giorn. Bot. Ital. 15: 189. (1883)
Source FNA vol. 11. Treatment authors: Michael A. Vincent, John M. Gillett†. FNA vol. 11.
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