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woolly-head clover
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balloon sack clover, cowbag clover, dwarf sack clover, poverty clover
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| Herbs perennial, 12–46 cm, sparsely hairy to densely villous, appressed-pubescent, or glabrate. |
Herbs annual, 1–30 cm, glabrous. |
erect or decumbent, branched. |
erect or decumbent, branched. |
palmate; stipules lanceolate or ovate, 1–5 cm, margins entire or denticulate, apex acuminate or acute; petiole 1–15 cm; petiolules 1 mm; leaflets 3, blades linear, linear-elliptic, elliptic, lanceolate, lanceolate-oblong, or obovate, 1–5 × 0.4–1.2 cm, base cuneate, veins fine or ± thickened, margins denticulate or serrulate, apex acuminate or acute, surfaces appressed-pubescent abaxially, glabrous, densely villous, or sparsely or densely pilose adaxially. |
palmate; stipules sheathing proximally, ovate-oblong to lanceolate, 0.4–1.3 cm, margins entire or serrulate, apex short triangular-cuspidate; petiole 1–5 cm; petiolules to 1 mm; leaflets 3, blades lanceolate, linear, obovate, or oblanceolate, 0.4–2.5 × 0.1–0.7 cm, base cuneate, veins moderately thickened abaxially, margins entire, serrate, or pinnately lobed, apex rounded, truncate, acute, or retuse, surfaces glabrous. |
terminal or axillary, 25–70-flowered, inverted or horizontal, ellipsoid or globose to subglobose, 2–3 × 1–3 cm, rachis prolonged beyond flowers; involucres absent. |
axillary or terminal, 3–15-flowered, globose or depressed-globose, 0.5–1.5 × 0.5–1.5 cm; involucres flattened, 3–13 mm, ± distinct or incised 1/2 their length, lobes 5–7, linear to elliptic or irregular, or vestigial, reduced to narrow ring, apex rounded to broad and irregularly erose, not split. |
bent distally, proximal to flowers, 3–18 cm. |
2–6 cm. |
reflexed, to 0.3 mm; bracteoles absent. |
straight, 0–0.5 mm; bracteoles absent. |
9–18 mm; calyx campanulate, curved in fruit, 4–11 mm, villous, veins 10, tube 1.5–2.5 mm, lobes unequal, 2–3 times tube, subulate, often strongly curved and twisted, plumose, orifice open; corolla white, cream, or purple, 8–16 mm, banner oblanceolate, 8–16 × 3–5 mm, apex rounded or retuse, recurved; ovaries pubescent distally. |
3–10 mm; calyx campanulate, 2–5 mm, glabrous, veins 5, tube 1–2.5 mm, lobes unequal, triangular-lanceolate or subulate, orifice open; corolla white or pink, 3–11 mm, inflated in fruit, banner broadly ovate, 4–7 × 3–8 mm, apex rounded, broad, acute, or retuse. |
ovoid, 2–3.5 mm. |
sessile or stipitate, ovoid, obovoid, or oblong, 2–4 mm. |
(1 or)2–4, yellow-brown to brown, mitten-shaped or subglobose, 1.2–2 mm, smooth. |
1–6, gray or tan, mottled, mitten-shaped, 1.2–1.6 mm, irregularly bumpy. |
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= 16. |
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w United States
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w North America; nw Mexico; c North America; South America (Chile, Peru)
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Subspecies 6 (6 in the flora). Trifolium eriocephalum was revised by J. M. Gillett (1971). The subspecies exhibit unusually complex distributional patterns, and overlapping morphological features sometimes make identifications problematic (M. Zohary and D. Heller 1984). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). The three varieties of Trifolium depauperatum are relatively easily distinguished by characteristics of the involucres and fruit stipes. Leaf morphology is not helpful in making the distinctions; in each variety, leaf margins can range from nearly entire or toothed or laciniate. In the phylogenetic treatment by N. W. Ellison et al. (2006), T. depauperatum is not allied with T. fucatum and other clovers with inflated corollas but rather with T. obtusiflorum, T. trichocalyx, and T. willdenovii (although with weak support). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Fabaceae > subfam. Faboideae > Trifolium |
Fabaceae > subfam. Faboideae > Trifolium |
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii |
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii |
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1. Ovules usually 2, rarely 1 or 3; calyx lobes usually green, rarely purple, straight, subequal, 3–4 timestube. | → 2 |
2. Herbs densely villous; leaflet blades of basal and median leaves elliptic or lanceolate-oblong; roots cylindric, thickened. | subsp. eriocephalum |
2. Herbs slightly villous, appressed-pubescent, or glabrate; leaflet blades of basal and median leaves usually narrowly lanceolate or linear, rarely narrowly elliptic; roots fusiform-tuberous. | subsp. cascadense |
1. Ovules 3 or 4; calyx lobes usually purple, contorted, abaxial lobes longer, lateral lobes 1–1.5 times tube. | → 3 |
3. Leaflet blades of basal leaves linear or linear-elliptic, apex acute, surfaces sparsely or densely hairy adaxially; roots cylindric, thickened, branched, often rhizomelike. | → 4 |
4. Blades of leaflets of median leaves linear or linear-elliptic; herbs slightly villous. | subsp. cusickii |
4. Blades of leaflets of median leaves broadly elliptic; herbs densely villous. | subsp. villiferum |
3. Leaflet blades of basal leaves obovate, apex rounded to retuse, surfaces usually glabrous adaxially, sometimes sparsely hairy; roots fusiform-tuberous. | → 5 |
5. Inflorescences 2.5 cm diam., significantly overtopping distal leaves; leaflet blades of median leaves broadly lanceolate, surfaces sparsely hairy or glabrous adaxially. | subsp. arcuatum |
5. Inflorescences 1 cm diam., scarcely overtopping distal leaves; leaflet blades of median leaves narrowly lanceolate or linear, surfaces densely villous adaxially. | subsp. martinii |
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1. Involucres reduced to narrow vestigial ring; inflorescences 1–1.5 cm diam.; flowers 7–10 mm; legumes ovoid, 3–4 mm, sessile. | var. depauperatum |
1. Involucres with evident bracts; inflorescences 0.5–1 cm diam.; flowers 3–8 mm; legumes oblong, ovoid, or obovoid, 2–4 mm, sessile or stipes 0.5–1 mm. | → 2 |
2. Involucral lobes broad, irregular, with broad, hyaline margins and reticulate veins; legumes oblong, 3–4 mm, ± sessile. | var. amplectens |
2. Involucral lobes oblong, often with narrow hyaline margins and raised, parallel veins; legumes ovoid or obovoid, 2–3 mm, stipes 0.5–1 mm. | var. truncatum |
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Lupinaster depauperatus |
| Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 313. (1838) |
Desvaux: J. Bot. Agric. 4: 69, plate 32, fig. 2. (1814) |
| FNA vol. 11. Treatment authors: Michael A. Vincent, John M. Gillett†. |
FNA vol. 11. |
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