woolly-head clover
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Santa Cruz clover
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Herbs perennial, 12–46 cm, sparsely hairy to densely villous, appressed-pubescent, or glabrate. |
Herbs annual, 5–40 cm, glabrous. |
erect or decumbent, branched. |
decumbent to ascending or erect, slightly branched. |
palmate; stipules lanceolate or ovate, 1–5 cm, margins entire or denticulate, apex acuminate or acute; petiole 1–15 cm; petiolules 1 mm; leaflets 3, blades linear, linear-elliptic, elliptic, lanceolate, lanceolate-oblong, or obovate, 1–5 × 0.4–1.2 cm, base cuneate, veins fine or ± thickened, margins denticulate or serrulate, apex acuminate or acute, surfaces appressed-pubescent abaxially, glabrous, densely villous, or sparsely or densely pilose adaxially. |
palmate; stipules whitish with prominent green veins, ovate, 0.3–0.5 cm, margins lacerate, apex acute to acuminate; petiole 1–2.4 cm; petiolules to 0.5 mm; leaflets 3, blades obovate to elliptic, 0.6–1.5 × 0.3–0.8 cm, base cuneate, veins ± thickened distally, margins denticulate, sometimes entire proximally, apex rounded, acute, or retuse, surfaces glabrous. |
terminal or axillary, 25–70-flowered, inverted or horizontal, ellipsoid or globose to subglobose, 2–3 × 1–3 cm, rachis prolonged beyond flowers; involucres absent. |
terminal or axillary, 15–20-flowered (early inflorescences hidden in stipules, subsessile, 2–5-flowered, flowers cleistogamous), subglobose, 0.5–0.8 × 0.6–0.8 cm; involucres flattened or shallowly bowl-shaped, 6–8 mm, when folded, nearly hiding calyces, glabrous or sparsely hairy, lobes 4 or 5, ± parallel-sided, 3 or 4-toothed. |
bent distally, proximal to flowers, 3–18 cm. |
1–3.5 cm. |
reflexed, to 0.3 mm; bracteoles absent. |
straight, to 0.5 mm; bracteoles absent. |
9–18 mm; calyx campanulate, curved in fruit, 4–11 mm, villous, veins 10, tube 1.5–2.5 mm, lobes unequal, 2–3 times tube, subulate, often strongly curved and twisted, plumose, orifice open; corolla white, cream, or purple, 8–16 mm, banner oblanceolate, 8–16 × 3–5 mm, apex rounded or retuse, recurved; ovaries pubescent distally. |
7–8 mm; calyx tubular, 4–5 mm, glabrous, veins 10, tube 2–2.5 mm, lobes subequal, triangular, margins conspicuously 2 or 3-toothed, apex aristate, orifice open; corolla pale pink or white, 6–7 mm, banner oblong, 5–7 × 1 mm, apex emarginate or erose. |
ovoid, 2–3.5 mm. |
ovoid, 2.5 mm. |
(1 or)2–4, yellow-brown to brown, mitten-shaped or subglobose, 1.2–2 mm, smooth. |
1, dark brown, slightly mottled, ovoid, 2 mm, smooth. |
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Flowering May–Jun. |
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Meadows, roadsides, grassy hillsides. |
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0–300 m. [0–1000 ft.] |
w United States
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CA |
Subspecies 6 (6 in the flora). Trifolium eriocephalum was revised by J. M. Gillett (1971). The subspecies exhibit unusually complex distributional patterns, and overlapping morphological features sometimes make identifications problematic (M. Zohary and D. Heller 1984). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Trifolium buckwestiorum, which ranges from Mendocino to Monterey counties, is unique among clovers in North America because of its aboveground cleistogamous, axillary flowers. The only other clover in North America that produces cleistogamous flowers is T. amphianthum, which produces its cleistogamous flowers at ground level and then pushes them into the substrate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Ovules usually 2, rarely 1 or 3; calyx lobes usually green, rarely purple, straight, subequal, 3–4 timestube. | → 2 |
2. Herbs densely villous; leaflet blades of basal and median leaves elliptic or lanceolate-oblong; roots cylindric, thickened. | subsp. eriocephalum |
2. Herbs slightly villous, appressed-pubescent, or glabrate; leaflet blades of basal and median leaves usually narrowly lanceolate or linear, rarely narrowly elliptic; roots fusiform-tuberous. | subsp. cascadense |
1. Ovules 3 or 4; calyx lobes usually purple, contorted, abaxial lobes longer, lateral lobes 1–1.5 times tube. | → 3 |
3. Leaflet blades of basal leaves linear or linear-elliptic, apex acute, surfaces sparsely or densely hairy adaxially; roots cylindric, thickened, branched, often rhizomelike. | → 4 |
4. Blades of leaflets of median leaves linear or linear-elliptic; herbs slightly villous. | subsp. cusickii |
4. Blades of leaflets of median leaves broadly elliptic; herbs densely villous. | subsp. villiferum |
3. Leaflet blades of basal leaves obovate, apex rounded to retuse, surfaces usually glabrous adaxially, sometimes sparsely hairy; roots fusiform-tuberous. | → 5 |
5. Inflorescences 2.5 cm diam., significantly overtopping distal leaves; leaflet blades of median leaves broadly lanceolate, surfaces sparsely hairy or glabrous adaxially. | subsp. arcuatum |
5. Inflorescences 1 cm diam., scarcely overtopping distal leaves; leaflet blades of median leaves narrowly lanceolate or linear, surfaces densely villous adaxially. | subsp. martinii |
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FNA vol. 11. |
FNA vol. 11. |
Fabaceae > subfam. Faboideae > Trifolium |
Fabaceae > subfam. Faboideae > Trifolium |
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii |
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii |
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Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 313. (1838) |
Isely: Madroño 39: 90, fig. 2. (1992) |
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