woolly-head clover
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Bolander's clover, parasol clover
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Herbs perennial, 12–46 cm, sparsely hairy to densely villous, appressed-pubescent, or glabrate. |
Herbs perennial, 15–30 cm, glabrous. |
erect or decumbent, branched. |
ascending, cespitose, branched. |
palmate; stipules lanceolate or ovate, 1–5 cm, margins entire or denticulate, apex acuminate or acute; petiole 1–15 cm; petiolules 1 mm; leaflets 3, blades linear, linear-elliptic, elliptic, lanceolate, lanceolate-oblong, or obovate, 1–5 × 0.4–1.2 cm, base cuneate, veins fine or ± thickened, margins denticulate or serrulate, apex acuminate or acute, surfaces appressed-pubescent abaxially, glabrous, densely villous, or sparsely or densely pilose adaxially. |
palmate; stipules ovate or lanceolate, 0.6–1.5 cm, margins entire, apex acute; petiole 1–7 cm; petiolules to 1 mm; leaflets 3, blades obcordate, obovate, or elliptic, 1–1.9 × 0.5–1.1 cm, base cuneate, veins fine to slightly thickened, margins setose, apex obtuse, rounded, or emarginate, apiculate, surfaces glabrous. |
terminal or axillary, 25–70-flowered, inverted or horizontal, ellipsoid or globose to subglobose, 2–3 × 1–3 cm, rachis prolonged beyond flowers; involucres absent. |
terminal, 12–30-flowered, ovoid, 1–2 × 1.5–3 cm, rachis prolonged beyond flowers, undivided or forked, often bearing sterile flower buds distally; involucres absent. |
bent distally, proximal to flowers, 3–18 cm. |
slender, bent distally, just below flowers, 5–20 cm. |
reflexed, to 0.3 mm; bracteoles absent. |
strongly reflexed, 1–1.2 mm; bracteoles minute, cuplike. |
9–18 mm; calyx campanulate, curved in fruit, 4–11 mm, villous, veins 10, tube 1.5–2.5 mm, lobes unequal, 2–3 times tube, subulate, often strongly curved and twisted, plumose, orifice open; corolla white, cream, or purple, 8–16 mm, banner oblanceolate, 8–16 × 3–5 mm, apex rounded or retuse, recurved; ovaries pubescent distally. |
12–14 mm; calyx deep violet, campanulate, gibbous, 3–5 mm, glabrous, veins 5, tube 1.5–2 mm, lobes unequal, triangular-subulate, orifice open; corolla lavender to purplish, 11–12 mm, banner curved, oblanceolate, 10–12 × 3–4 mm, apex narrow, rounded, slightly emarginate. |
ovoid, 2–3.5 mm. |
ellipsoid, 3–4 mm. |
(1 or)2–4, yellow-brown to brown, mitten-shaped or subglobose, 1.2–2 mm, smooth. |
1 or 2, brown, ellipsoid, 1.2–1.5 mm, smooth. |
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Flowering Jun–Aug. |
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Meadows. |
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2000–2300 m. [6600–7500 ft.] |
w United States
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CA |
Subspecies 6 (6 in the flora). Trifolium eriocephalum was revised by J. M. Gillett (1971). The subspecies exhibit unusually complex distributional patterns, and overlapping morphological features sometimes make identifications problematic (M. Zohary and D. Heller 1984). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Trifolium bolanderi is known from the Sierra Nevada in California and may be vulnerable to shifts in climate patterns. Genetic diversity of populations of the species (based on allozyme patterns) was found to be relatively high; two genetically distinct groups of populations are known (R. G. Denton 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Ovules usually 2, rarely 1 or 3; calyx lobes usually green, rarely purple, straight, subequal, 3–4 timestube. | → 2 |
2. Herbs densely villous; leaflet blades of basal and median leaves elliptic or lanceolate-oblong; roots cylindric, thickened. | subsp. eriocephalum |
2. Herbs slightly villous, appressed-pubescent, or glabrate; leaflet blades of basal and median leaves usually narrowly lanceolate or linear, rarely narrowly elliptic; roots fusiform-tuberous. | subsp. cascadense |
1. Ovules 3 or 4; calyx lobes usually purple, contorted, abaxial lobes longer, lateral lobes 1–1.5 times tube. | → 3 |
3. Leaflet blades of basal leaves linear or linear-elliptic, apex acute, surfaces sparsely or densely hairy adaxially; roots cylindric, thickened, branched, often rhizomelike. | → 4 |
4. Blades of leaflets of median leaves linear or linear-elliptic; herbs slightly villous. | subsp. cusickii |
4. Blades of leaflets of median leaves broadly elliptic; herbs densely villous. | subsp. villiferum |
3. Leaflet blades of basal leaves obovate, apex rounded to retuse, surfaces usually glabrous adaxially, sometimes sparsely hairy; roots fusiform-tuberous. | → 5 |
5. Inflorescences 2.5 cm diam., significantly overtopping distal leaves; leaflet blades of median leaves broadly lanceolate, surfaces sparsely hairy or glabrous adaxially. | subsp. arcuatum |
5. Inflorescences 1 cm diam., scarcely overtopping distal leaves; leaflet blades of median leaves narrowly lanceolate or linear, surfaces densely villous adaxially. | subsp. martinii |
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FNA vol. 11. |
FNA vol. 11. |
Fabaceae > subfam. Faboideae > Trifolium |
Fabaceae > subfam. Faboideae > Trifolium |
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii |
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii |
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Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 313. (1838) |
A. Gray: Proc. Amer. Acad. Arts 7: 335. (1868) |
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