woolly-head clover
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Beckwith's clover
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Herbs perennial, 12–46 cm, sparsely hairy to densely villous, appressed-pubescent, or glabrate. |
Herbs perennial, 5–50 cm, glabrous. |
erect or decumbent, branched. |
erect or ascending, unbranched or several stems clumped from crown. |
palmate; stipules lanceolate or ovate, 1–5 cm, margins entire or denticulate, apex acuminate or acute; petiole 1–15 cm; petiolules 1 mm; leaflets 3, blades linear, linear-elliptic, elliptic, lanceolate, lanceolate-oblong, or obovate, 1–5 × 0.4–1.2 cm, base cuneate, veins fine or ± thickened, margins denticulate or serrulate, apex acuminate or acute, surfaces appressed-pubescent abaxially, glabrous, densely villous, or sparsely or densely pilose adaxially. |
palmate; stipules narrowly ovate, lanceolate, or linear-lanceolate, 1–2 cm, margins entire or toothed, apex acute; petiole 0.5–20 cm; petiolules 1 mm; leaflets 3, blades usually rhombic, oblong, or elliptic, sometimes ovate, 2.5–5 × 0.6–2 cm, base cuneate, veins slightly thickened, margins setose-serrulate, apex obtuse or retuse, surfaces glaucous abaxially, glabrous. |
terminal or axillary, 25–70-flowered, inverted or horizontal, ellipsoid or globose to subglobose, 2–3 × 1–3 cm, rachis prolonged beyond flowers; involucres absent. |
terminal or axillary, 30–70+-flowered, globose or ovoid becoming subglobose, 1.5–4 × 1.8–3.3 cm, rachis prolonged beyond flowers, undivided or forked, often bearing sterile flower buds distally; involucres absent. |
bent distally, proximal to flowers, 3–18 cm. |
straight, thick, 5–25 cm. |
reflexed, to 0.3 mm; bracteoles absent. |
strongly reflexed in fruit, 1–1.5 mm; bracteoles minute. |
9–18 mm; calyx campanulate, curved in fruit, 4–11 mm, villous, veins 10, tube 1.5–2.5 mm, lobes unequal, 2–3 times tube, subulate, often strongly curved and twisted, plumose, orifice open; corolla white, cream, or purple, 8–16 mm, banner oblanceolate, 8–16 × 3–5 mm, apex rounded or retuse, recurved; ovaries pubescent distally. |
11–14 mm, sweetly fragrant; calyx campanulate, gibbous, 4–6 mm, glabrous, veins 5, tube 2–3 mm, lobes unequal, narrowly triangular or subulate, orifice open; corolla light purplish, sometimes with pink tips, 10–13 mm, banner straight, obovate to broadly elliptic, 13–18 × 6–8 mm, apex rounded or retuse, apiculate. |
ovoid, 2–3.5 mm. |
oblong, 5 mm. |
(1 or)2–4, yellow-brown to brown, mitten-shaped or subglobose, 1.2–2 mm, smooth. |
2–4, reddish brown, flattened globose, 1.5–2 mm, smooth. |
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= 48. |
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Flowering May–Jul. |
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Moist, grassy meadows along streams. |
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1200–2000 m. [3900–6600 ft.] |
w United States
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CA; ID; MT; NV; OR; SD; UT; WA
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Subspecies 6 (6 in the flora). Trifolium eriocephalum was revised by J. M. Gillett (1971). The subspecies exhibit unusually complex distributional patterns, and overlapping morphological features sometimes make identifications problematic (M. Zohary and D. Heller 1984). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Disjunct populations of Trifolium beckwithii found in South Dakota are over 1200 km east of the nearest populations in Montana. The South Dakota populations appear to have been long-distance introductions from populations in northern California, based on molecular data (M. R. Duvall et al. 1999). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Ovules usually 2, rarely 1 or 3; calyx lobes usually green, rarely purple, straight, subequal, 3–4 timestube. | → 2 |
2. Herbs densely villous; leaflet blades of basal and median leaves elliptic or lanceolate-oblong; roots cylindric, thickened. | subsp. eriocephalum |
2. Herbs slightly villous, appressed-pubescent, or glabrate; leaflet blades of basal and median leaves usually narrowly lanceolate or linear, rarely narrowly elliptic; roots fusiform-tuberous. | subsp. cascadense |
1. Ovules 3 or 4; calyx lobes usually purple, contorted, abaxial lobes longer, lateral lobes 1–1.5 times tube. | → 3 |
3. Leaflet blades of basal leaves linear or linear-elliptic, apex acute, surfaces sparsely or densely hairy adaxially; roots cylindric, thickened, branched, often rhizomelike. | → 4 |
4. Blades of leaflets of median leaves linear or linear-elliptic; herbs slightly villous. | subsp. cusickii |
4. Blades of leaflets of median leaves broadly elliptic; herbs densely villous. | subsp. villiferum |
3. Leaflet blades of basal leaves obovate, apex rounded to retuse, surfaces usually glabrous adaxially, sometimes sparsely hairy; roots fusiform-tuberous. | → 5 |
5. Inflorescences 2.5 cm diam., significantly overtopping distal leaves; leaflet blades of median leaves broadly lanceolate, surfaces sparsely hairy or glabrous adaxially. | subsp. arcuatum |
5. Inflorescences 1 cm diam., scarcely overtopping distal leaves; leaflet blades of median leaves narrowly lanceolate or linear, surfaces densely villous adaxially. | subsp. martinii |
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FNA vol. 11. |
FNA vol. 11. |
Fabaceae > subfam. Faboideae > Trifolium |
Fabaceae > subfam. Faboideae > Trifolium |
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii |
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii |
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Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 313. (1838) |
Brewer ex S. Watson: Proc. Amer. Acad. Arts 11: 128. (1876) |
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