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woolly-head clover

Egyptian clover

Habit Herbs perennial, 12–46 cm, sparsely hairy to densely villous, appressed-pubescent, or glabrate. Herbs annual, 20–60 cm, appressed-pubescent.
Stems

erect or decumbent, branched.

erect, branched.

Leaves

palmate;

stipules lanceolate or ovate, 1–5 cm, margins entire or denticulate, apex acuminate or acute;

petiole 1–15 cm;

petiolules 1 mm;

leaflets 3, blades linear, linear-elliptic, elliptic, lanceolate, lanceolate-oblong, or obovate, 1–5 × 0.4–1.2 cm, base cuneate, veins fine or ± thickened, margins denticulate or serrulate, apex acuminate or acute, surfaces appressed-pubescent abaxially, glabrous, densely villous, or sparsely or densely pilose adaxially.

palmate;

stipules lanceolate, 1–2 cm, margins denticulate distally, plumose-ciliate, apex caudate;

petiole 1–8 cm;

petiolules 1+ mm;

leaflets 3, blades broadly elliptic to oblong, 1–5 × 0.5–1.5 cm, base cuneate, veins fine, margins denticulate, apex usually mucronate, sometimes retuse, surfaces hairy.

Inflorescences

terminal or axillary, 25–70-flowered, inverted or horizontal, ellipsoid or globose to subglobose, 2–3 × 1–3 cm, rachis prolonged beyond flowers;

involucres absent.

terminal, 25–100-flowered, ovoid, elongate in fruit, 1–2.5 × 1–1.5 cm;

involucres minute or absent.

Peduncles

bent distally, proximal to flowers, 3–18 cm.

0.5–9 cm.

Pedicels

reflexed, to 0.3 mm;

bracteoles absent.

absent;

bracteoles absent.

Flowers

9–18 mm;

calyx campanulate, curved in fruit, 4–11 mm, villous, veins 10, tube 1.5–2.5 mm, lobes unequal, 2–3 times tube, subulate, often strongly curved and twisted, plumose, orifice open;

corolla white, cream, or purple, 8–16 mm, banner oblanceolate, 8–16 × 3–5 mm, apex rounded or retuse, recurved;

ovaries pubescent distally.

13–16 mm;

calyx cylindric (campanulate in fruit), 6–8 mm, appressed-hairy, veins 10, tube 1–2 mm, lobes erect to slightly spreading, equal or abaxial 2 slightly longer, narrowly triangular, orifice open, hairy;

corolla cream or yellowish white, 13–16 mm, banner narrowly spatulate, 13–16 × 3–4 mm, apex obtuse.

Legumes

ovoid, 2–3.5 mm.

ellipsoid, leathery distally, transversely dehiscent, 2.2–2.5 mm.

Seeds

(1 or)2–4, yellow-brown to brown, mitten-shaped or subglobose, 1.2–2 mm, smooth.

1, yellow, ovoid, 2–2.2 mm, smooth, glossy.

2n

= 16.

Trifolium eriocephalum

Trifolium alexandrinum

Phenology Flowering Apr–May.
Habitat Disturbed areas, fields.
Elevation 0–50 m. [0–160 ft.]
Distribution
from USDA
w United States
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; w Asia (Syria) [Introduced in North America; introduced also in e Asia (e China), n Africa, Australia]
[BONAP county map]
Discussion

Subspecies 6 (6 in the flora).

Trifolium eriocephalum was revised by J. M. Gillett (1971). The subspecies exhibit unusually complex distributional patterns, and overlapping morphological features sometimes make identifications problematic (M. Zohary and D. Heller 1984).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trifolium alexandrinum (known also as berseem, from its Arabic name) has been cultivated in the Mediterranean region for millennia and appears to have been introduced into Egypt in the sixth century (P. B. Kennedy and W. W. Mackie 1925). The origin of T. alexandrinum was long shrouded in mystery, because it appeared that the progenitors of the species were possibly extinct (W. E. Knight 1985b). Examination of similar species using molecular data gave indications that T. berytheum Boissier & C. I. Blanche and T. salmoneum Mouterde might be the primary ancestors from which domestication occurred through artificial selection in Syria (A. Badr et al. 2008).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ovules usually 2, rarely 1 or 3; calyx lobes usually green, rarely purple, straight, subequal, 3–4 timestube.
→ 2
2. Herbs densely villous; leaflet blades of basal and median leaves elliptic or lanceolate-oblong; roots cylindric, thickened.
subsp. eriocephalum
2. Herbs slightly villous, appressed-pubescent, or glabrate; leaflet blades of basal and med­ian leaves usually narrowly lanceolate or lin­ear, rarely narrowly elliptic; roots fusiform-tuberous.
subsp. cascadense
1. Ovules 3 or 4; calyx lobes usually purple, contorted, abaxial lobes longer, lateral lobes 1–1.5 times tube.
→ 3
3. Leaflet blades of basal leaves linear or linear-elliptic, apex acute, surfaces sparsely or densely hairy adaxially; roots cylindric, thickened, branched, often rhizomelike.
→ 4
4. Blades of leaflets of median leaves linear or linear-elliptic; herbs slightly villous.
subsp. cusickii
4. Blades of leaflets of median leaves broadly elliptic; herbs densely villous.
subsp. villiferum
3. Leaflet blades of basal leaves obovate, apex rounded to retuse, surfaces usually glabrous adaxially, sometimes sparsely hairy; roots fusiform-tuberous.
→ 5
5. Inflorescences 2.5 cm diam., significantly overtopping distal leaves; leaflet blades of median leaves broadly lanceolate, surfaces sparsely hairy or glabrous adaxially.
subsp. arcuatum
5. Inflorescences 1 cm diam., scarcely over­topping distal leaves; leaflet blades of median leaves narrowly lanceolate or linear, surfaces densely villous adaxially.
subsp. martinii
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Trifolium Fabaceae > subfam. Faboideae > Trifolium
Sibling taxa
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
T. albopurpureum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
Subordinate taxa
T. eriocephalum subsp. arcuatum, T. eriocephalum subsp. cascadense, T. eriocephalum subsp. cusickii, T. eriocephalum subsp. eriocephalum, T. eriocephalum subsp. martinii, T. eriocephalum subsp. villiferum
Name authority Nuttall in J. Torrey and A. Gray: Fl. N. Amer. 1: 313. (1838) Linnaeus: Cent. Pl. I, 25. (1755)
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