Trifolium columbinum |
Trifolium carolinianum |
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olive clover |
Carolina clover |
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Habit | Herbs annual, 8–30 cm, canescent. | Herbs annual, 5–30 cm, pilose. |
Stems | erect, branched from base and distally, or unbranched. |
prostrate, ascending, or erect, branched. |
Leaves | palmate; stipules ovate, 0.4–1.2 cm, margins entire, ciliate, apex often purple, acuminate, setaceous; petiole 0.5–8 cm; petiolules 0.5–1 mm; leaflets 3, blades obovate or elliptic, 0.5–3 × 0.3–1.2 cm, base cuneate, veins obscure or slightly thickened, margins dentate, apex rounded or emarginate, surfaces pilose. |
palmate; stipules ovate, 0.6–1 cm, margins entire or serrate, apex acuminate; petiole 2–5 cm; petiolules 1 mm; leaflets 3, blades obovate, 0.6–1.4 × 0.3–1.1 cm, base cuneate, veins moderately thickened, margins denticulate, apex rounded, sometimes retuse, surfaces glabrous except on veins abaxially. |
Inflorescences | terminal or axillary, 25–50-flowered, ovoid-ellipsoid, 1–2.5 × 1.2–2.5 cm; involucres absent. |
axillary or terminal, 5–40-flowered, globose, 1–2 × 1–2 cm; involucres minute. |
Peduncles | 3–11 cm. |
± geniculate proximal to flowers, inflorescence appearing sideways or inverted, 3.5–10 cm. |
Pedicels | absent; bracteoles cuplike, to 0.5 mm. |
reflexed, 2–3 mm; bracteoles ovate or lanceolate, 0.5 mm. |
Flowers | 6–10 mm; calyx tubular, 8–14 mm, densely pubescent, veins 20–30, tube 1–2 mm, lobes nearly equal, appearing rigidly erect, linear-setaceous, plumose, sinuses acute, orifice open; corolla sometimes absent, red-violet, often with white tips, 4–6 mm, banner ovate-oblong, 7–8 × 2–3 mm, apex broadly rounded, apiculate. |
4.5–7 mm; calyx usually pink to magenta, campanulate, bilabiate, 3–4 mm, pilose, veins 5, slightly branched in lobes, tube 0.8–1 mm, lobes unequal, abaxial 3 lanceolate or linear, 0.8–1 mm, villous, adaxial 2 broadly triangular, 2.5–3 mm, orifice open; corolla creamy white, often pink-tinged, darkening in age, 4–4.5 mm, banner broadly ovate-rhombic, 4–4.5 × 3.8–4 mm, apex rounded, slightly erose. |
Legumes | obovoid, 3–3.5 mm. |
oblong, 3–4 mm. |
Seeds | 1, yellow-brown, ellipsoid, 2–2.5 mm, smooth. |
2–4, yellow-brown to brown, globose to mitten-shaped, 1.1–1.6 mm, rugose. |
2n | = 16. |
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Trifolium columbinum |
Trifolium carolinianum |
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Phenology | Flowering Apr–May. | Flowering Apr–Jun. |
Habitat | Grassy slopes, meadows, heavy adobe/clay soils, oak woodland foothills. | Sandy soils in dry fields, lawns, open woods, disturbed ground, rocky slopes. |
Elevation | 0–1400 m. (0–4600 ft.) | 0–400 m. (0–1300 ft.) |
Distribution |
CA |
AL; AR; DE; FL; GA; KS; LA; MO; MS; NC; OK; PA; SC; TX; VA
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Discussion | D. Isely (1998) stated that Central Valley populations of Trifolium columbinum appeared to be extinct and populations outside that region (and here recognized as the same taxon) were similar in calyx and corolla proportions. It is rather widespread in California, ranging from Humboldt County southward to Santa Barbara County, and eastward into counties ranging from Shasta southward to Inyo, Kern, and Placer counties. Trifolium albopurpureum var. olivaceum (Greene) Isely and T. columbinum var. olivaceum (Greene) Jepson are illegitimate names that pertain here (D. J. Keil 2019). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Trifolium arvense Walter (1788), a later homonym of T. arvense Linnaeus (1753), was considered a synonym of T. carolinianum by M. Zohary and D. Heller (1984). Trifolium repens Walter (1788, not Linnaeus 1753) is likely also a synonym of T. carolinianum. The record of the species from Vermont is a waif. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | T. albopurpureum var. columbinum, T. columbinum var. argillorum, T. olivaceum, T. olivaceum var. columbinum, T. olivaceum var. griseum | Amoria caroliniana, T. oxypetalum, T. umbellatum |
Name authority | Greene: Pittonia 1: 4. (1887) | Michaux: Fl. Bor.-Amer. 2: 58. (1803) |
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