Trifolium ciliolatum |
Trifolium cyathiferum |
|
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foothill clover, tree clover |
bowl clover, cup clover |
|
Habit | Herbs annual, 5–50 cm, glabrous or sparsely hairy. | Herbs annual, 5–50 cm, glabrous. |
Stems | erect, branched. |
erect, branched. |
Leaves | palmate; stipules ovate-lanceolate, 1–1.5 cm, margins entire, sometimes ciliate, apex acuminate; petiole 1–13 cm; petiolules 0.5 mm; leaflets 3, blades elliptic to oblong or obovate, 0.8–3.5 × 0.5–1.5 cm, base cuneate, veins thickened, margins serrate proximally, obscurely denticulate distally, apex usually rounded or retuse, rarely acute, surfaces glabrous. |
palmate; stipules ovate to lanceolate, 0.6–1.2 cm, margins entire, toothed, or lacerate, apex acute or aristate; petiole 1–10 cm; petiolules to 0.5 mm; leaflets 3, blades obovate, elliptic, or oblanceolate, 0.5–2.5 × 0.3–1.5 cm, base cuneate, veins prominent, green, margins finely toothed, apex rounded, blunt, or retuse, surfaces glabrous. |
Inflorescences | axillary or terminal, 10–30-flowered, ovoid to subglobose, 0.7–2.2 × 0.5–2 cm; involucres a narrow rim, 0.5 mm, membranous, dentate. |
axillary or terminal, 1–35-flowered, subglobose or ovoid, 0.5–1.8 × 0.5–2 cm; involucres broadly bowl-shaped, 4–22 mm, shallowly incised, lobes 3–15, toothed, broad, acute. |
Peduncles | 2.5–12 cm. |
1–8 cm. |
Pedicels | erect becoming reflexed, 0.5–6 mm; bracteoles linear or cup-shaped, to 1 mm. |
straight, 0.1–0.2 mm; bracteoles absent. |
Flowers | 6–13 mm; calyx broadly campanulate, 5–11 mm, glabrous, veins 10, tube 1–5 mm, lobes unequal, elliptic to linear, margins hyaline, dentate or pectinate, ciliate, sinuses narrow, orifice open; corolla white, pink, or purple, 5–13 mm, banner broadly ovate, 6–13 × 4–7 mm, apex rounded, apiculate. |
7–15 mm; calyx campanulate, inflated in fruit, 6–13 mm, glabrous, veins 13–20, tube 3–7 mm, lobes unequal, strongly oblique, broadly triangular, apex setaceous, adaxial unbranched, abaxial and lateral conspicuously 2- or 3-fid, segments glabrous, sometimes flattened, orifice open; corolla usually creamy white or rose to pink, sometimes whitish with pinkish tips, 6–13 mm, banner obovate or elliptic, proximally inflated in fruit, distally narrowed into twisted tip, 6–13 × 3–6 mm, apex rounded to broadly acute. |
Legumes | short-stipitate, ovoid, 5–10 mm. |
ellipsoid, 2.5–3 mm. |
Seeds | 1 or 2, brown, mottled, ovoid, 2.5–3 mm, smooth. |
1 or 2, yellow-brown, ovoid, 0.4–0.6 mm, smooth. |
2n | = 16. |
= 16. |
Trifolium ciliolatum |
Trifolium cyathiferum |
|
Phenology | Flowering Apr–Jun. | Flowering Apr–Oct. |
Habitat | Oak-pine chaparral, meadows, roadsides. | Wet meadows, roadsides, fields. |
Elevation | 0–1500 m. (0–4900 ft.) | 0–2700 m. (0–8900 ft.) |
Distribution |
CA; OR; WA; Mexico (Baja California)
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AK; CA; ID; MT; NV; OR; WA; BC; YT [Introduced in Asia (China, Japan)]
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Discussion | Trifolium ciliolatum is relatively widespread in California and is found in scattered sites in Baja California, Oregon, and Washington. Trifolium ciliatum Nuttall (1848), which pertains here, is a later homonym of T. ciliatum E. D. Clarke (1813). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Trifolium cyathiferum may be of hybrid origin; phylogenetic studies indicate it has the same trnL intron sequence as T. buckwestiorum, T. polyodon, and T. variegatum and nrDNA and combined analyses place it in a clade with other species (T. barbigerum, T. fucatum, T. jokerstii, and T. physanthum Hooker & Arnott) that have inflated fruiting corollas (N. W. Ellison et al. 2006). The single record of Trifolium cyathiferum labeled as from Utah is likely mislabeled (S. L. Welsh et al. 2008). Populations in Yukon may be introductions. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | Fabaceae > subfam. Faboideae > Trifolium | Fabaceae > subfam. Faboideae > Trifolium |
Sibling taxa | ||
Synonyms | T. ciliatum var. discolor | |
Name authority | Bentham: Pl. Hartw., 304. (1849) | Lindley: Bot. Reg. 13: sub plate 1070. (1827) |
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