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Bolander's clover, parasol clover

olive clover

Habit Herbs perennial, 15–30 cm, glabrous. Herbs annual, 8–30 cm, canescent.
Stems

ascending, cespitose, branched.

erect, branched from base and distally, or unbranched.

Leaves

palmate;

stipules ovate or lanceolate, 0.6–1.5 cm, margins entire, apex acute;

petiole 1–7 cm;

petiolules to 1 mm;

leaflets 3, blades obcordate, obovate, or elliptic, 1–1.9 × 0.5–1.1 cm, base cuneate, veins fine to slightly thickened, margins setose, apex obtuse, rounded, or emarginate, apiculate, surfaces glabrous.

palmate;

stipules ovate, 0.4–1.2 cm, margins entire, ciliate, apex often purple, acuminate, setaceous;

petiole 0.5–8 cm;

petiolules 0.5–1 mm;

leaflets 3, blades obovate or elliptic, 0.5–3 × 0.3–1.2 cm, base cuneate, veins obscure or slightly thickened, margins dentate, apex rounded or emarginate, surfaces pilose.

Inflorescences

terminal, 12–30-flowered, ovoid, 1–2 × 1.5–3 cm, rachis prolonged beyond flowers, undivided or forked, often bearing sterile flower buds distally;

involucres absent.

terminal or axillary, 25–50-flowered, ovoid-ellipsoid, 1–2.5 × 1.2–2.5 cm;

involucres absent.

Peduncles

slender, bent distally, just below flowers, 5–20 cm.

3–11 cm.

Pedicels

strongly reflexed, 1–1.2 mm;

bracteoles minute, cuplike.

absent;

bracteoles cuplike, to 0.5 mm.

Flowers

12–14 mm;

calyx deep violet, campanulate, gibbous, 3–5 mm, glabrous, veins 5, tube 1.5–2 mm, lobes unequal, triangular-subulate, orifice open;

corolla lavender to purplish, 11–12 mm, banner curved, oblanceolate, 10–12 × 3–4 mm, apex narrow, rounded, slightly emarginate.

6–10 mm;

calyx tubular, 8–14 mm, densely pubescent, veins 20–30, tube 1–2 mm, lobes nearly equal, appearing rigidly erect, linear-setaceous, plumose, sinuses acute, orifice open;

corolla sometimes absent, red-violet, often with white tips, 4–6 mm, banner ovate-oblong, 7–8 × 2–3 mm, apex broadly rounded, apiculate.

Legumes

ellipsoid, 3–4 mm.

obovoid, 3–3.5 mm.

Seeds

1 or 2, brown, ellipsoid, 1.2–1.5 mm, smooth.

1, yellow-brown, ellipsoid, 2–2.5 mm, smooth.

Trifolium bolanderi

Trifolium columbinum

Phenology Flowering Jun–Aug. Flowering Apr–May.
Habitat Meadows. Grassy slopes, meadows, heavy adobe/clay soils, oak woodland foothills.
Elevation 2000–2300 m. (6600–7500 ft.) 0–1400 m. (0–4600 ft.)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
Discussion

Trifolium bolanderi is known from the Sierra Nevada in California and may be vulnerable to shifts in climate patterns. Genetic diversity of populations of the species (based on allozyme patterns) was found to be relatively high; two genetically distinct groups of populations are known (R. G. Denton 2002).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. Isely (1998) stated that Central Valley populations of Trifolium columbinum appeared to be extinct and populations outside that region (and here recognized as the same taxon) were similar in calyx and corolla proportions. It is rather widespread in California, ranging from Humboldt County southward to Santa Barbara County, and eastward into counties ranging from Shasta southward to Inyo, Kern, and Placer counties.

Trifolium albopurpureum var. olivaceum (Greene) Isely and T. columbinum var. olivaceum (Greene) Jepson are illegitimate names that pertain here (D. J. Keil 2019).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Trifolium Fabaceae > subfam. Faboideae > Trifolium
Sibling taxa
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. columbinum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
T. albopurpureum, T. alexandrinum, T. amoenum, T. amphianthum, T. andersonii, T. andinum, T. angustifolium, T. appendiculatum, T. arvense, T. attenuatum, T. aureum, T. barbigerum, T. barnebyi, T. beckwithii, T. bejariense, T. bifidum, T. bolanderi, T. brandegeei, T. breweri, T. buckwestiorum, T. calcaricum, T. campestre, T. carolinianum, T. cernuum, T. ciliolatum, T. cyathiferum, T. dasyphyllum, T. dedeckerae, T. depauperatum, T. dichotomum, T. douglasii, T. dubium, T. echinatum, T. eriocephalum, T. fragiferum, T. friscanum, T. fucatum, T. glomeratum, T. gracilentum, T. grayi, T. gymnocarpon, T. haydenii, T. hirtum, T. howellii, T. hybridum, T. hydrophilum, T. incarnatum, T. jokerstii, T. kentuckiense, T. kingii, T. lappaceum, T. latifolium, T. leibergii, T. lemmonii, T. longipes, T. lupinaster, T. macilentum, T. macraei, T. macrocephalum, T. medium, T. microcephalum, T. microdon, T. monanthum, T. mucronatum, T. nanum, T. nigrescens, T. obtusiflorum, T. oliganthum, T. ornithopodioides, T. owyheense, T. palmeri, T. parryi, T. pinetorum, T. piorkowskii, T. plumosum, T. polyodon, T. pratense, T. productum, T. reflexum, T. repens, T. resupinatum, T. retusum, T. rollinsii, T. siskiyouense, T. sonorense, T. stoloniferum, T. striatum, T. subterraneum, T. suffocatum, T. thompsonii, T. tomentosum, T. trichocalyx, T. variegatum, T. vesiculosum, T. virginicum, T. willdenovii, T. wormskioldii
Synonyms T. albopurpureum var. columbinum, T. columbinum var. argillorum, T. olivaceum, T. olivaceum var. columbinum, T. olivaceum var. griseum
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 335. (1868) Greene: Pittonia 1: 4. (1887)
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