Trifolium andersonii |
Trifolium polyodon |
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Anderson's clover, fiveleaf clover |
Pacific grove clover, woods or Pacific grove clover |
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Habit | Herbs perennial, 5–12 cm, canescent. | Herbs annual, 10–60 cm, glabrous. | ||||||||
Stems | erect-ascending, cespitose, mat-forming, branched; from deep taproot-like crown. |
decumbent or ascending, branched. |
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Leaves | palmate; stipules ovate or lanceolate, 0.5–1.5 cm, margins entire, apex long-acuminate; petiole 1–5 cm; petiolules to 0.5 mm; leaflets (3–)5, blades obovate, 0.5–2.2 × 0.2–1.2 cm, base cuneate, veins obscured by pubescence, margins entire, apex rounded or acute, apiculate, surfaces canescent. |
palmate; stipules lanceolate to ovate, 0.4–1.8 cm, margins lacerate, apex acute to acuminate; petiole 0.5–6 cm; petiolules to 1 mm; leaflets 3, blades broadly elliptic to broadly obovate, 0.4–2.5 × 0.4–1.5 cm, base cuneate, veins moderately thickened, margins dentate-serrulate, apex rounded or truncate, often retuse, surfaces glabrous. |
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Inflorescences | axillary, 10–20-flowered, depressed-globose, 1–2 × 1–2.3 cm; involucres formed of connate, narrow, membranous bracts, to 0.5 mm. |
axillary, 10–25-flowered, subglobose or globose, 1–1.8 × 1–1.8 cm; involucres flattened or bowl-shaped, 4–7 mm, incised 1/2 their length, when folded, not hiding flowers except proximally, lobes 5–12, dentate-lacerate, spinulose. |
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Peduncles | 1–14 cm. |
1–2 cm. |
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Pedicels | absent; bracteoles absent. |
straight, to 1 mm; bracteoles absent. |
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Flowers | 10–17 mm; calyx campanulate, 9–11 mm, pilose, veins 10, tube 4–5 mm, lobes equal, subulate, plumose, orifice open; corolla white or pink, 10–17 mm, banner ovate-oblong, 12–19 × 3–5 mm, apex rounded, retuse. |
8–10 mm; calyx campanulate-tubular, not slit between adaxial lobes, 5–7 mm, glabrous, veins 20, tube 1.9–2.4 mm, lobes unequal, 3-fid or laciniate, often appearing 7+-lobed, orifice open; corolla pink or pale purple, 8–9 mm, banner elliptic, 8–9 × 2–3 mm, apex retuse. |
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Legumes | ellipsoid, 4–5 mm. |
ellipsoid, 3.5–4 mm. |
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Seeds | 1 (or 2), tan or brown, irregularly ovoid, 1.6–2 mm, smooth, glossy. |
2, dark brown, mottled, ovoid or reniform, 1.7–1.9 mm, smooth. |
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Trifolium andersonii |
Trifolium polyodon |
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Phenology | Flowering Apr–Jun. | |||||||||
Habitat | Along streams, moist meadows. | |||||||||
Elevation | 0–150 m. (0–500 ft.) | |||||||||
Distribution |
w United States
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CA |
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Discussion | Subspecies 3 (3 in the flora). Trifolium andersonii encompasses three moderately well separable subspecies, with subsp. andersonii the more northern form in the range of the species, subsp. monoense in the southwestern portion of the range, and subsp. beatleyae in the eastern portion of the range (J. M. Gillett 1972). Intermediates between the latter two (and overlap in their ranges) make differentiation between them somewhat problematic, which prompted R. C. Barneby (1989) to combine them as var. beatleyae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The taxonomic status of Trifolium polyodon, known only from Monterey County, has long been debated. W. L. Jepson (1936) considered it a variety of T. tridentatum Lindley (= T. willdenovii). The first to consider it as a variety of T. variegatum was J. S. Martin (1943), who never formally published a new combination; an invalid combination was made by M. Zohary and D. Heller (1984). D. Isely (1998) included it in his interpretation of T. variegatum as phase 5 of that species; he speculated that T. polyodon might have originated as a hybrid of T. variegatum and T. willdenovii. Molecular studies (N. W. Ellison et al. 2006) showed that T. polyodon is closely related to T. variegatum but is distinct; it also appears to be related to T. cyathiferum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 11. | FNA vol. 11. | ||||||||
Parent taxa | Fabaceae > subfam. Faboideae > Trifolium | Fabaceae > subfam. Faboideae > Trifolium | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | T. tridentatum var. polyodon | |||||||||
Name authority | A. Gray: Proc. Amer. Acad. Arts 6: 522. (1865) | Greene: Pittonia 3: 215. (1897) | ||||||||
Web links |