Trifolium amoenum |
Trifolium polyodon |
|
---|---|---|
showy Indian clover, two-fork clover |
Pacific grove clover, woods or Pacific grove clover |
|
Habit | Herbs annual, 45–65 cm, canescent. | Herbs annual, 10–60 cm, glabrous. |
Stems | erect, branched from base and distally, or unbranched. |
decumbent or ascending, branched. |
Leaves | palmate; stipules ovate-oblong, 0.5–1.8 cm, margins entire, toothed, or irregularly lobed, apex acuminate; petiole 0.5–10 cm; petiolules to 1 mm; leaflets 3, blades broadly obovate or elliptic, 1.7–3.3 × 1.1–2 cm, base cuneate, veins fine, margins entire or denticulate, apex rounded, obtuse, or retuse, surfaces pilose, abaxial less so. |
palmate; stipules lanceolate to ovate, 0.4–1.8 cm, margins lacerate, apex acute to acuminate; petiole 0.5–6 cm; petiolules to 1 mm; leaflets 3, blades broadly elliptic to broadly obovate, 0.4–2.5 × 0.4–1.5 cm, base cuneate, veins moderately thickened, margins dentate-serrulate, apex rounded or truncate, often retuse, surfaces glabrous. |
Inflorescences | terminal or axillary, 30–50-flowered, bluntly conic, ellipsoid, globose, or subglobose, 1.5–3 × 1.5–2.5 cm; involucres absent. |
axillary, 10–25-flowered, subglobose or globose, 1–1.8 × 1–1.8 cm; involucres flattened or bowl-shaped, 4–7 mm, incised 1/2 their length, when folded, not hiding flowers except proximally, lobes 5–12, dentate-lacerate, spinulose. |
Peduncles | 5–15 cm. |
1–2 cm. |
Pedicels | absent; bracteoles cuplike, to 0.5 mm. |
straight, to 1 mm; bracteoles absent. |
Flowers | 13–16 mm; calyx tubular, 9–12 mm, pubescent, veins 20–30, tube 3 mm, lobes nearly equal, appearing rigidly erect, linear-setaceous, plumose, sinuses acute, orifice open; corolla white to pink with purple tips, 12–16 mm, banner ovate-oblong, 12–15 × 4 mm, apex broadly rounded, emarginate. |
8–10 mm; calyx campanulate-tubular, not slit between adaxial lobes, 5–7 mm, glabrous, veins 20, tube 1.9–2.4 mm, lobes unequal, 3-fid or laciniate, often appearing 7+-lobed, orifice open; corolla pink or pale purple, 8–9 mm, banner elliptic, 8–9 × 2–3 mm, apex retuse. |
Legumes | obovoid, 4 mm. |
ellipsoid, 3.5–4 mm. |
Seeds | 1, dark brown, ellipsoid, 2–3 mm, smooth. |
2, dark brown, mottled, ovoid or reniform, 1.7–1.9 mm, smooth. |
Trifolium amoenum |
Trifolium polyodon |
|
Phenology | Flowering Apr–May. | Flowering Apr–Jun. |
Habitat | Grassy slopes, swales, clay soils. | Along streams, moist meadows. |
Elevation | 0–100 m. (0–300 ft.) | 0–150 m. (0–500 ft.) |
Distribution |
CA
|
CA |
Discussion | Trifolium amoenum was considered extinct (J. P. Smith Jr. 1984) but was rediscovered in 1993 (P. G. Connors 1994). Specimens of T. amoenum are known from Marin, Napa, San Mateo, Solano, and Sonoma counties; it appears to be extant in only single populations in each of Marin and Solano counties. Allozyme studies of the two known populations revealed fixed genetic differences between them (E. E. Knapp and Connors 1999). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The taxonomic status of Trifolium polyodon, known only from Monterey County, has long been debated. W. L. Jepson (1936) considered it a variety of T. tridentatum Lindley (= T. willdenovii). The first to consider it as a variety of T. variegatum was J. S. Martin (1943), who never formally published a new combination; an invalid combination was made by M. Zohary and D. Heller (1984). D. Isely (1998) included it in his interpretation of T. variegatum as phase 5 of that species; he speculated that T. polyodon might have originated as a hybrid of T. variegatum and T. willdenovii. Molecular studies (N. W. Ellison et al. 2006) showed that T. polyodon is closely related to T. variegatum but is distinct; it also appears to be related to T. cyathiferum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | T. tridentatum var. polyodon | |
Name authority | Greene: Fl. Francisc., 27. (1891) | Greene: Pittonia 3: 215. (1897) |
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