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marsh, marsh St. John's wort

greater marsh St. Johnswort

Habit Herbs, perennial, rhizomatous, glabrous, with glandular canals, lacunae, or dots containing essential oils (pale) in various parts and, sometimes, reddish to purplish gland dots containing hypericin on stems and leaves. Herbs erect, to 10 dm, sometimes with ascending branches in distal 1/2+.
Stems

internodes with 2 or 4 raised lines at first, then terete (not lined).

internodes 4-lined at first, then terete.

Leaves

sessile, subsessile, or petiolate;

blade relatively broad, venation pinnate, tertiary veins densely reticulate, glands punctiform, pale (records of black gland dots are probably all due to fungal attack), intramarginal and laminar.

petiolate, petiole 2–15 mm;

blade narrowly elliptic to narrowly oblong, 30–150 × 10–35 mm, base cuneate, apex rounded, gland dots laminar and intramarginal.

Inflorescences

terminal, sometimes also axillary, cymose, 2–15-flowered, or solitary flower, branching dichasial;

bracts and bracteoles relatively small.

interrupted spiciform-cylindric, 3-flowered from terminal node, sometimes with subsessile flowers or pedunculate triads and/or flowering branches from to 8 proximal nodes.

Flowers

tubular or campanulate at first, expanding to stellate for short time each day;

sepals persistent, 5, distinct or almost so, margins not glandular-ciliate;

petals deciduous, 5, partly imbricate or contorted, pink or flesh-colored, sometimes green-tinged;

stamens persistent, 9, in 3 fascicles, each with 3 stamens;

filaments of each fascicle 1/5–1/2+ connate;

anthers yellow, isodiametric to oblate or shortly oblong, with amber gland on connective;

staminode fascicles 3, alternating with stamen fascicles;

ovary 3-merous;

placentation axile;

ovules relatively numerous on each placenta;

styles distinct, spreading.

7–10 mm diam.;

sepals narrowly oblong to narrowly elliptic or lanceolate, 3–5 × 1–1.5 mm, apex obtuse to rounded;

petals narrowly obovate, 5–7 mm;

stamen fascicles 3–5 mm;

filaments 1/2+ connate;

styles 0.8–1.2 mm.

Capsules

3-valved, with glandular vittae.

cylindric, 7–12 × 3–4 mm, apex subacute.

Seeds

narrowly cylindric, carinate;

testa reticulate-foveolate.

0.8–1.1 mm.

x

= 19.

Triadenum

Triadenum walteri

Phenology Flowering late summer (Jul–Sep).
Habitat Swampy or marshy ground in woods, pond or lake margins, on fallen logs
Elevation 0–400 m (0–1300 ft)
Distribution
from USDA
e North America; Asia (e China, India [Assam], Japan, Korea, e Siberia, Taiwan)
from FNA
AL; AR; DE; FL; GA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; OH; OK; SC; TN; TX; VA; WV
[WildflowerSearch map]
Discussion

Nomenclatural complexities and confusions associated with Triadenum were reviewed by N. K. B. Robson (1977). B. R. Ruhfel et al. (2011) concluded from molecular studies that Triadenum is part of Hypericum. Robson (2012) gave reasons why Triadenum is generically distinct.

Species 6 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The petiolate leaves usually distinguish Triadenum walteri from the other three North American species of Triadenum (see note under T. tubulosum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves petiolate.
T. walteri
1. Leaves sessile or (distal) subsessile
→ 2
2. Leaves: gland dots intramarginal.
T. tubulosum
2. Leaves: gland dots laminar and intramarginal
→ 3
3. Sepals 4–7(–8) mm, apex acute to acuminate; styles 2–3.5 mm.
T. virginicum
3. Sepals 2.5–5 mm, apex usually obtuse to rounded, rarely acute; styles 0.5–1.5 mm.
T. fraseri
Source FNA vol. 6, p. 102. FNA vol. 6, p. 105.
Parent taxa Hypericaceae Hypericaceae > Triadenum
Sibling taxa
T. fraseri, T. tubulosum, T. virginicum
Subordinate taxa
T. fraseri, T. tubulosum, T. virginicum, T. walteri
Synonyms Gardenia, Hypericum section Elodea Hypericum walteri, Elodes floribunda, E. petiolata, Gardenia petiolata, Hypericum paludosum, H. tubulosum var. walteri, T. petiolatum
Name authority Rafinesque: Fl. Tellur. 3: 78. (1837) (J. F. Gmelin) Gleason: Phytologia 2: 289. (1947)
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