Triadenum |
Triadenum tubulosum |
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marsh, marsh St. John's wort |
lesser marsh St. Johnswort |
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Habit | Herbs, perennial, rhizomatous, glabrous, with glandular canals, lacunae, or dots containing essential oils (pale) in various parts and, sometimes, reddish to purplish gland dots containing hypericin on stems and leaves. | Herbs erect, to 10 dm, sometimes branching from near base, sometimes with ascending branches in distal 1/2+. | ||||||||||||
Stems | internodes with 2 or 4 raised lines at first, then terete (not lined). |
internodes 2-lined at first, then terete. |
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Leaves | sessile, subsessile, or petiolate; blade relatively broad, venation pinnate, tertiary veins densely reticulate, glands punctiform, pale (records of black gland dots are probably all due to fungal attack), intramarginal and laminar. |
sessile or (distal) subsessile, rarely amplexicaul; blade narrowly oblong or elliptic to oblanceolate, 50–120(–150) × 10–40(–50) mm, base rounded to truncate or subcordate, apex rounded or rounded-apiculate to retuse, gland dots intramarginal. |
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Inflorescences | terminal, sometimes also axillary, cymose, 2–15-flowered, or solitary flower, branching dichasial; bracts and bracteoles relatively small. |
spiciform-cylindric, 3–7-flowered from terminal node, with sessile or pedunculate inflorescences and flowering branches from to 4 proximal nodes. |
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Flowers | tubular or campanulate at first, expanding to stellate for short time each day; sepals persistent, 5, distinct or almost so, margins not glandular-ciliate; petals deciduous, 5, partly imbricate or contorted, pink or flesh-colored, sometimes green-tinged; stamens persistent, 9, in 3 fascicles, each with 3 stamens; filaments of each fascicle 1/5–1/2+ connate; anthers yellow, isodiametric to oblate or shortly oblong, with amber gland on connective; staminode fascicles 3, alternating with stamen fascicles; ovary 3-merous; placentation axile; ovules relatively numerous on each placenta; styles distinct, spreading. |
15 mm diam.; sepals narrowly oblong, 4–7 × 1–1.5 mm, apex acute to obtuse; petals elliptic-obovate, 5–8 mm; stamen fascicles 4–7 mm; filaments 1/2+ connate; styles 0.8–1.5 mm. |
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Capsules | 3-valved, with glandular vittae. |
cylindric, 8–12 × 3.5–4 mm, apex obtuse. |
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Seeds | narrowly cylindric, carinate; testa reticulate-foveolate. |
0.8–1.2 mm. |
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x | = 19. |
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Triadenum |
Triadenum tubulosum |
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Phenology | Flowering late summer–early fall (Aug–Sep). | |||||||||||||
Habitat | Swampy or marshy ground in woods | |||||||||||||
Elevation | 0–500 m (0–1600 ft) | |||||||||||||
Distribution |
e North America; Asia (e China, India [Assam], Japan, Korea, e Siberia, Taiwan) |
AL; AR; FL; GA; IL; IN; KY; LA; MO; MS; NC; OH; SC; TN; TX; VA
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Discussion | Nomenclatural complexities and confusions associated with Triadenum were reviewed by N. K. B. Robson (1977). B. R. Ruhfel et al. (2011) concluded from molecular studies that Triadenum is part of Hypericum. Robson (2012) gave reasons why Triadenum is generically distinct. Species 6 (4 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The absence of laminar glands in leaves is a more reliable characteristic for differentiating Triadenum tubulosum from T. walteri than sessile versus petiolate leaves, which are not always quite sessile towards the inflorescence in T. tubulosum, or sepals, which can be obtuse in both. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 102. | FNA vol. 6, p. 104. | ||||||||||||
Parent taxa | ||||||||||||||
Sibling taxa | ||||||||||||||
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Synonyms | Gardenia, Hypericum section Elodea | Hypericum tubulosum, Elodes drummondii, E. pauciflora, E. tubulosa, H. petiolatum var. tubulosum, H. walteri var. tubulosum, T. longifolium | ||||||||||||
Name authority | Rafinesque: Fl. Tellur. 3: 78. (1837) | (Walter) Gleason: Phytologia 2: 289. (1947) | ||||||||||||
Web links |