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long-neck moss, trematodon moss

trematodon moss

Leaves

ovate-lanceolate to long-subulate from an ovate or obovate base, serrulate at apex;

costa percurrent or ending before the apex.

usually contorted when dry, lanceolate to broadly subulate from an ovate or oblong sheathing base, 1.5–4 mm, apex acute to narrowly obtuse, margins entire or with a few teeth apically;

costa subpercurrect to excurrent as a subula;

distal laminal cells subquadrate or short-rectangular.

Seta

long, 10–30 mm.

elongate, 0.5–1.5 cm.

Capsule

inclined, curved;

neck 2–3 times as long as urn when dry, long-cylindric, strumose at base;

peristome teeth 2-fid or irregularly perforate, not fragile, commonly persistent;

annulus compound, revoluble.

long-exserted, cylindric with neck strongly differentiated, conic to long-cylindric;

peristome absent or present, of 16 simple, forked or perforate teeth, vertically barred on external surface;

operculum differentiated, obliquely long-rostrate.

Calyptra

cucullate.

Spores

medium-sized, 20–30 µm, papillose.

Perichaetial

leaves distinctly larger than the cauline, convolute-sheathing.

Trematodon longicollis

Trematodon

Phenology Capsules mature spring–summer.
Habitat Soil, sand
Elevation low to moderate elevations
Distribution
from FNA
AL; AR; FL; GA; KY; LA; MD; MS; NC; NJ; OH; OK; PA; SC; TX; VA; Mexico; Central America; South America; West Indies (Cuba, Puerto Rico); Asia (China, Japan, Papua New Guinea); Pacific Islands (New Caledonia)
[WildflowerSearch map]
Worldwide; mainly temperate areas
Discussion

Trematodon longicollis is much like T. ambiguus but differs in the relative lengths of capsule urn and neck when dry, and is clearly more southern in distribution. The peristome may occasionally adhere to the operculum on dehiscence and the capsule may then falsely appear gymnostomous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 25 (6 in the flora).

Trematodon is clearly in need of revision, with more than 80 correct binomials extant but only about 25 species commonly accepted as well characterized. It clearly intergrades with Bruchia. Within the genus, only T. ambiguus and T. longicollis have strumose capsules.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Seta short, 1-4 mm; peristome teeth undivided, annulus simple
→ 2
1. Seta long, (4-)10-30 mm; peristome teeth split, or perforate basally or apically, occasionally entire, annulus usually compound, revoluble
→ 3
2. Seta 1.5-2 mm; capsule neck short, barely as long as the urn, obovate; peristome fragile, often lost with dehiscence of operculum; leaves abruptly subulate, entire at apex.
T. montanus
2. Seta 1-4 mm; capsule neck slightly longer than the urn, tapering-conic; peristome persistent; leaves long-lanceolate, weakly denticulate apically.
T. boasii
3. Capsule neck strumose
→ 4
3. Capsule neck not strumose
→ 5
4. Capsule neck about 2-3 times the length of urn.
T. longicollis
4. Capsule neck about the length of the urn.
T. ambiguus
5. Leaves broad, ovate-lanceolate; capsule somewhat curved, peristome teeth entire or perforate near apex.
T. brevicollis
5. Leaves narrow, mostly long-lanceolate; capsule straight, peristome teeth perforate medially and often split near apex.
T. laetevirens
Source FNA vol. 27, p. 438. FNA vol. 27, p. 437.
Parent taxa Bruchiaceae > Trematodon Bruchiaceae
Sibling taxa
T. ambiguus, T. boasii, T. brevicollis, T. laetevirens, T. montanus
Subordinate taxa
T. ambiguus, T. boasii, T. brevicollis, T. laetevirens, T. longicollis, T. montanus
Synonyms T. ambiguus var. longicollis
Name authority Michaux: Fl. Bor.-Amer. 2: 289. (1803) Michaux: Fl. Bor.-Amer. 2: 289. 1803 ,
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