The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

long-neck moss, trematodon moss

Habit Plants minute or merely small, gregarious to densely tufted.
Stems

erect, simple or branching, with a central strand.

Leaves

ovate-lanceolate to long-subulate from an ovate or obovate base, serrulate at apex;

costa percurrent or ending before the apex.

mostly lanceolate or subulate, straight or somewhat curved, base oblong to ovate; in several rows;

margins plane, entire or serrulate distally;

costa single, well developed, subpercurrent to shortly excurrent as a sometimes roughened or denticulate subula, in section poorly differentiated or with 1 row of guide cells and 2 (sub-)stereid bands, adaxial band much reduced;

lamina cells smooth or abaxially papillose;

basal cells broader, narrower towards the margins, those of basal angles not differentiated or forming a marginal border;

distal cells short- to long-rectangular, walls firm.

Seta

long, 10–30 mm.

short to elongate, usually yellow or brown;

capsules immersed to exerted, cylindric or obovate with a distinct inflated-tapering to elongate neck, erect to curved; cleistocarpous, gymnostomous, or peristomate;

annulus, when present, usually of 2–3 rows of larger cells, commonly revoluble;

peristome, when present, single, of 16 simple, forked, or perforate teeth;

operculum,when present, obliquely long-rostrate.

Sexual condition

autoicous or paroicous [dioicous];

perigonia axillary or on short branches adjacent to perichaetia or basal on the plant;

perichaetial leaves usually differentiated, longer and somewhat sheathing.

Capsule

inclined, curved;

neck 2–3 times as long as urn when dry, long-cylindric, strumose at base;

peristome teeth 2-fid or irregularly perforate, not fragile, commonly persistent;

annulus compound, revoluble.

Calyptra

cucullate or mitrate.

Spores

often large, spheric to ovoid or weakly reniform, finely to coarsely papillose, spiculate, reticulate, or pitted.

Specialized

asexual reproduction not known.

Trematodon longicollis

Bruchiaceae

Phenology Capsules mature spring–summer.
Habitat Soil, sand
Elevation low to moderate elevations
Distribution
from FNA
AL; AR; FL; GA; KY; LA; MD; MS; NC; NJ; OH; OK; PA; SC; TX; VA; Mexico; Central America; South America; West Indies (Cuba, Puerto Rico); Asia (China, Japan, Papua New Guinea); Pacific Islands (New Caledonia)
[WildflowerSearch map]
Cosmopolitan; greatest occurrence in temperate regions
Discussion

Trematodon longicollis is much like T. ambiguus but differs in the relative lengths of capsule urn and neck when dry, and is clearly more southern in distribution. The peristome may occasionally adhere to the operculum on dehiscence and the capsule may then falsely appear gymnostomous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 4, species ca. 140 (2 genera, 16 species in the flora).

Species of Bruchiaceae commonly occur on soil, often as colonizers. The family was well-characterized by W. R. Buck (1979), who recognized four genera and suggested a transitional position between Dicranaceae and Ditrichaceae. The two genera represented in the flora area are quite different but share the salient well-developed capsule neck.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Capsule cleistocarpic, immersed to short-exserted, neck conic to ovate.
Bruchia
1. Capsule stegocarpic, long-exserted, neck conic to long-cylindric.
Trematodon
Source FNA vol. 27, p. 438. FNA vol. 27, p. 433. Author: Richard H. Zander.
Parent taxa Bruchiaceae > Trematodon
Sibling taxa
T. ambiguus, T. boasii, T. brevicollis, T. laetevirens, T. montanus
Subordinate taxa
Bruchia, Trematodon
Synonyms T. ambiguus var. longicollis
Name authority Michaux: Fl. Bor.-Amer. 2: 289. (1803) Schimper
Web links