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spike bur grass, spike burr grass

bur grass

Habit Plants annual. Plants annual or perennial; cespitose.
Culms

(2)3.5-45 cm.

(2)5-65 cm, herbaceous, usually rooting at the lower nodes;

nodes and internodes glabrous.

Leaves

cauline;

sheaths open, usually shorter than the internodes, mostly glabrous but long-ciliate at the edges of the collar;

ligules membranous, truncate, ciliate;

blades usually flat, margins ciliate.

Panicles

(1)2-13 cm long, (3)4-8 mm wide;

rachises pubescent;

branches (0.5)0.7-2.7 mm, pubescent, with 2(3) spikelets, axes occasionally extending past the distal spikelets;

proximal internodes 0.2-0.6(0.7) mm, shorter than the second internodes.

Inflorescences

terminal, exceeding the upper leaves, narrow, cylindrical panicles;

branches 0.5-5 mm, resembling burs, with 2-5 spikelets;

disarticulation at the base of the branches.

Spikelets

crowded, attached individually to the branches, with 1 floret;

proximal spikelet(s) bisexual, larger than the distal spikelet(s);

terminal spikelets often sterile.

Glumes

unequal;

lower glumes absent or minute, veinless, membranous;

upper glumes usually exceeding the florets, 5-7-veined, with 5-7 longitudinal rows of straight or uncinate spinelike projections;

lemmas 3-veined;

paleas 2-veined, hyaline, membranous, x = 10.

Caryopses

(0.9)1.2-2 mm long, 0.4-0.8 mm wide.

Ligules

0.5-1 mm;

blades (0.5)0.7-8.5 cm long, 1.2-5 mm wide, glabrous.

Proximal

spikelets (1.8)2^.3 mm;

second spikelets (0.8)1-3.9 mm, sometimes sterile.

Lower

glumes 0.1-0.6 mm, membranous, minutely pubescent;

upper glumes 1.8-4.3 mm, minutely pubescent, 5-veined, rarely with 1-2 additional veins adjacent to the midvein;

glume projections (4)6-14, in 5 rows, (0.2)0.3-1 mm, uncinate;

lemmas (1.5)1.8-3.1 mm, sparsely pubescent on the back, midveins occasionally excurrent to 0.6 mm;

paleas (1.3)1.5-2.4 mm;

anthers 3, 0.4-0.6 mm, yellow, occasionally purple-or green-tinged.

2n

= 20.

Tragus berteronianus

Tragus

Distribution
from FNA
AZ; MA; ME; NM; NY; SC; TX; VA; HI; PR; Virgin Islands
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; MA; MD; ME; NC; NJ; NM; NY; PA; SC; TX; VA; HI; PR; Virgin Islands
[BONAP county map]
Discussion

Tragus berteronianus is native to Africa and Asia, and is now established in Arizona, New Mexico, and Texas. It was collected in Maine, Massachusetts, New York, and Virginia in the nineteenth century, and Virginia in 1959.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Tragus has seven species, all of which are native to the tropics and subtropics of the Eastern Hemisphere; four have been introduced into the Flora region. The genus is easily recognized by the spinelike projections on the upper glumes. The number of veins in the glume should be determined by examining the adaxial surface, where they appear as green lines.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Upper glumes with 5 longitudinal rows of spinelike projections, 5-veined.
→ 2
2. Proximal internodes of the primary branches not longer than the second internode
T. berteronianus
2. Proximal internodes of the primary branches 2-3 (or more) times longer than the second internode
T. australianus
1. Upper glumes with (5)6-7 longitudinal rows of spinelike projections, 7-veined.
→ 3
3. Panicle branches with 2 (rarely 3) spikelets; proximal spikelets on the branches 3-3.5 mm long
T. heptaneuron
3. Panicle branches with 3-5 (rarely 2) spikelets; proximal spikelets on the branches 3.8-6.6 mm long
T. racemosus
Source FNA vol. 25, p. 280. FNA vol. 25, p. 278. Author: J.K. Wipff;.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Tragus Poaceae > subfam. Chloridoideae > tribe Cynodonteae
Sibling taxa
T. australianus, T. heptaneuron, T. racemosus
Subordinate taxa
T. australianus, T. berteronianus, T. heptaneuron, T. racemosus
Name authority Schult. Haller
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